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Regulation of peptidoglycan synthesis and remodelling

Nat Rev Microbiol. 2020 Aug;18(8):446-460. doi: 10.1038/s41579-020-0366-3. Epub 2020 May 18.

Abstract

Bacteria surround their cell membrane with a net-like peptidoglycan layer, called sacculus, to protect the cell from bursting and maintain its cell shape. Sacculus growth during elongation and cell division is mediated by dynamic and transient multiprotein complexes, the elongasome and divisome, respectively. In this Review we present our current understanding of how peptidoglycan synthases are regulated by multiple and specific interactions with cell morphogenesis proteins that are linked to a dynamic cytoskeletal protein, either the actin-like MreB or the tubulin-like FtsZ. Several peptidoglycan synthases and hydrolases require activation by outer-membrane-anchored lipoproteins. We also discuss how bacteria achieve robust cell wall growth under different conditions and stresses by maintaining multiple peptidoglycan enzymes and regulators as well as different peptidoglycan growth mechanisms, and we present the emerging role of LD-transpeptidases in peptidoglycan remodelling.

Publication types

  • Research Support, Non-U.S. Gov't
  • Review

MeSH terms

  • Bacteria / metabolism
  • Bacterial Proteins / metabolism*
  • Cell Membrane / metabolism
  • Cell Wall / metabolism
  • Cytoskeletal Proteins / metabolism
  • Peptidoglycan / metabolism*

Substances

  • Bacterial Proteins
  • Cytoskeletal Proteins
  • Peptidoglycan