Reliable data transport in wireless sensor networks is a multifaceted problem influenced by the physical, MAC, network, and transport layers. Because sensor networks are subject to strict resource constraints and are deployed by single organizations, they encourage revisiting traditional layering and are less bound by standardized placement of services such as reliability. This paper presents analysis and experiments resulting in specific recommendations for implementing reliable data transport in sensor nets. To explore reliability at the transport layer, we present RMST (Reliable Multi-Segment Transport), a new transport layer for Directed Diffusion. RMST provides guaranteed delivery and fragmentation/reassembly for applications that require them. RMST is a selective NACK-based protocol that can be configured for in-network caching and repair.