FIGURE 12. Bivariate plots with 95% confidence regions for the first two axes derived from scores... more FIGURE 12. Bivariate plots with 95% confidence regions for the first two axes derived from scores of DFA analyses for male (A) and female (B) colour patterns of Atractus paraguayensis.
FIGURE 6. Sulcate (left) and asulcate (right) sides of the hemipenis of (A) Atractus francoi (IBS... more FIGURE 6. Sulcate (left) and asulcate (right) sides of the hemipenis of (A) Atractus francoi (IBSP 72654), (B) A. guentheri (MNRJ 6710), (C) A. maculatus (MNRJ 14197), (D) A. potschi (MZUEFS 682), (E) A. pantostictus (MNRJ 13195), and (F) A. paraguaguayensis (MNRJ 10123). Scale = 5 mm.
FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. fr... more FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46).
FIGURE 10. General view in life of Atractus paraguayensis. Specimens from Chapecó, state of Santa... more FIGURE 10. General view in life of Atractus paraguayensis. Specimens from Chapecó, state of Santa Catarina (A, C, and D), photos by F.M. D'Agostini. Specimens from Segredo Hydroeletric Power Plant, Reserva do Iguaçú, state of Paraná (B and F), photos by M.V. Segalla. Specimen from Passo Fundo River, Nanoai, state of Rio Grande do Sul (E), photo by N.R. Albuquerque.
FIGURE 4. Geographical distribution of Atractus caete, A. guentheri, A. maculatus, A. potschi, an... more FIGURE 4. Geographical distribution of Atractus caete, A. guentheri, A. maculatus, A. potschi, and A. ronnie.
<i>Atractus zebrinus</i> (Jan, 1862) Figs. 18B, 19, 20 <i>Rabdosoma zebrinum<... more <i>Atractus zebrinus</i> (Jan, 1862) Figs. 18B, 19, 20 <i>Rabdosoma zebrinum</i> Jan, 1862; Arch. Zool. Anat. Fis. 2:15. <i>Rhabdosoma maculatus</i> – Günther, 1858; Catalogue of the Colubrine Snakes in the British Museum:421. (part.). <i>Atractus maculatus</i> – Boulenger, 1894; Catalogue of the Snakes in the British Museum 2:306. (part.). <i>Atractus guentheri</i> – Amaral, 1978; Serpentes do Brasil: Iconografia Colorida:114. <i>Atractus</i> cf. <i>guentheri</i> – Lema, 1994; Comun. Mus. Ciênc. Tecnol. PUCRS, Sér. Zool. 7:105. (part.). <i>Atractus maculatus</i> – Moura-Leite, Morato &amp; Bérnils, 1996; Herp. Rev. 27:216. <i>Atractus zebrinus</i> – Fernandes, Freire &amp; Puorto, 2000; Bol. Mus. Nac. Nov. Sér. Zool. 419:4. <b>Holotype:</b> Specimen without defined locality, originally housed at Museo Civico di Storia Naturale di Milano (Jan 1862), believed to have been destroyed during the Second World War (S. Scali pers. comm.). <b>Diagnosis:</b> <i>Atractus zebrinus</i> is distinguished from all congeners by the following combination of characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) loreal moderate; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) generally seven infralabials, first four contacting chinshields; (7) eight to ten maxillary teeth; (8) generally three gular scale rows; (9) generally three preventrals; (10) 149–170 ventrals in females, 136–153 in males; (11) 15–27 subcaudals in females, 20–32 in males; (12) dorsum creamish red with irregular transverse black blotches with white borders in adults; (13) venter immaculate creamish white; (14) moderate body size, with females reaching 600 mm SVL and males 457 mm SVL; (15) tail short in females (6.8–11.7% SVL), moderate to long (10.0–16.5% SVL) in males; (16) hemipenis slightly or moderate bilobed, semicapitate, and semicalyculate. <b>Comparisons:</b> Among all congeners, <i>A. zebrinus</i> shares 17 dorsal scale rows, seven upper and lower labials, first four infralabiais contacting chinshi [...]
<i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007 Figs. 13C, 15A... more <i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007 Figs. 13C, 15A <i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007; Copeia 2007:789. <i>Atractus ronnie</i> – Loebmann, Ribeiro, Sales &amp; Almeida, 2009; Biotemas 22:170. <b>Holotype:</b> Adult female, MNRJ 14194, from Serra de Baturité (04°10'S, 38°55'W, ca. 800 m), municipality of Pacoti in the state of Ceará, Brazil, collected on 10 April 1998 by D. M. Borges-Nojosa (specimen examined). <b>Paratypes:</b> Twenty-four specimens, all from Serra do Baturité, state of Ceará: municipality of Pacoti (CHUFC 1396, MNRJ 14195), collected on 17 July 1989 by L. W. Lima-Verde; (CHUFC 2646), 18 December 1997 by D. M. Borges-Nojosa; (CHUFC 2648), 11 January 1998 by D. M. Borges-Nojosa; (CHUFC 2658), 09 December 2005 by D. M. Borges-Nojosa, J. C. L. Melo, and M. J. B. Leite, locality of Sítio Olho d`água dos Tangarás; (CHUFC 2481, 3500), 23 February 1989 by D. M. Borges-Nojosa, locality of Monguba; (CHUFC 3502), 03 April 1990 by D. M. Borges-Nojosa, locality of Sítio São José; (MNRJ 14196–97), 16 February 1999 and 10 April 1998, respectively by D. M. Borges-Nojosa, locality of Granja; (CHUFC 2641, 2647), 02 November 1997 by D. M. Borges-Nojosa, locality of Sítio Pau do Alho; (CHUFC 25980), 28 April 2005 by W. C. M. Luz, locality of Sítio Xangrilá; (CHUFC 2652–24), 2005 December by W. C. M. Luz, locality of Cidade Pacoti; (CHUFC 2675–76, 2678), between 17 February and 02 March 2006 by W. C. M. Luz, locality of Cidade Pacoti; (CHUFC 2733), 04 April 1990 by D. M. Borges-Nojosa, district of Santana. Municipality of Baturité: (CHUFC 2578), 11 February 2005 by D. M. Borges-Nojosa, P. Cascon, and J. C. L. Melo, locality of Sítio Escuro. Municipality of Mulungu: (CHUFC 2645), 1991 by D. M. Borges-Nojosa and S. M. Cornélio. Municipality of Guaramiranga: (CHUFC 2649), 15 August 1998 by D. M. Borges-Nojosa, locality of Linha da Serra; (CHUFC 2651), 6 June 2005 by D. M. Borges-Nojosa, P. Cascon, and J. C. L. Melo, locality of Sítio Guaramiranga. <b>Diagno [...]
FIGURE 13. General view of original Sordelli's plate of the Atractus multicinctus. Modified f... more FIGURE 13. General view of original Sordelli's plate of the Atractus multicinctus. Modified from Jan & Sordelli (1865).
FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B)... more FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B) detailed view of lateral expansion of the hemipenis of Atractus iridesnces, (C) sulcate and asulcate sides of the hemipenis of Atractus typhon sp.nov. (ICN 10901). Scale = 5mm.
FIGURE 10. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) ... more FIGURE 10. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) views of head of the lectotype of Tricheilostoma macrolepis (ZMB 1434).
FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostom... more FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostoma brevissimum (MCZ 38950).
FIGURE 3. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) v... more FIGURE 3. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) views of head of the lectotype of Trilepida macrolepis (ZMB 1434). Photos by F. Tillack (ZMB).
(2002) FERNANDES et al. Boletim do Museu nacional. Read by researchers in: 100% Biological Scienc... more (2002) FERNANDES et al. Boletim do Museu nacional. Read by researchers in: 100% Biological Sciences. The taxonomic status of the lowland species of the Liophis cobella group is revised, with the recognition of four species: L. cobella (Linnaeus, 1758), L. breviceps Cope, 1860 ...
The taxonomic status of Atractus sanctaemartae and A. nebularis is revised on the basis of quanti... more The taxonomic status of Atractus sanctaemartae and A. nebularis is revised on the basis of quantitative and qualitative analyses of morphological characters (meristics, morphometrics, colour pattern and hemipenis). Characters used previously for diagnosingAtractus nebularis from A. sanctaemartae are demystified, and the synonymy of these species is proposed. Colour pattern, meristic, morphometrical and scale ornamentation differences originally employed in recognizing both taxa are explained as sexually dimorphic characters, which hold no geographic or phylogenetic basis. Additionally, a new species closely related to Atractus sanctaemartae is described from the Atlantic coast of Colombia. It is distinguished from A. sanctaemartae mainly by having two gular scale rows, slender (62% of body diameter) and acuminate head, snout acuminate in lateral view, dorsum uniformly reddish brown and venter cream with a series of rhomboidal blotches arranged linearly on the lateral portion of the ...
We report on the morphological variation (meristic, morphometric, pholidosis, and color pattern c... more We report on the morphological variation (meristic, morphometric, pholidosis, and color pattern characters) and sexual dimorphism of Trilepida macrolepis. We also took the opportunity to redescribe the holotype of Rena affinis in an effort to evaluate its taxonomic status. Based on the congruence of characters from external morphology, we transfer Stenostoma affine to the genus Trilepida. We also discuss the Andean specimens previously refereed to Trilepida affinis and Trilepida macrolepis and provide an identification key for the genus Trilepida.
FIGURE 12. Bivariate plots with 95% confidence regions for the first two axes derived from scores... more FIGURE 12. Bivariate plots with 95% confidence regions for the first two axes derived from scores of DFA analyses for male (A) and female (B) colour patterns of Atractus paraguayensis.
FIGURE 6. Sulcate (left) and asulcate (right) sides of the hemipenis of (A) Atractus francoi (IBS... more FIGURE 6. Sulcate (left) and asulcate (right) sides of the hemipenis of (A) Atractus francoi (IBSP 72654), (B) A. guentheri (MNRJ 6710), (C) A. maculatus (MNRJ 14197), (D) A. potschi (MZUEFS 682), (E) A. pantostictus (MNRJ 13195), and (F) A. paraguaguayensis (MNRJ 10123). Scale = 5 mm.
FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. fr... more FIGURE 2. General view of the holotypes of (A) Atractus caete sp. nov. (MNRJ 16936), (B, C) A. francoi sp. nov. (MNRJ 17537), (D) A. guentheri (BMNH 1946.1.1.76), and (E) A. maculatus (BMNH 1946.1.6.46).
FIGURE 10. General view in life of Atractus paraguayensis. Specimens from Chapecó, state of Santa... more FIGURE 10. General view in life of Atractus paraguayensis. Specimens from Chapecó, state of Santa Catarina (A, C, and D), photos by F.M. D'Agostini. Specimens from Segredo Hydroeletric Power Plant, Reserva do Iguaçú, state of Paraná (B and F), photos by M.V. Segalla. Specimen from Passo Fundo River, Nanoai, state of Rio Grande do Sul (E), photo by N.R. Albuquerque.
FIGURE 4. Geographical distribution of Atractus caete, A. guentheri, A. maculatus, A. potschi, an... more FIGURE 4. Geographical distribution of Atractus caete, A. guentheri, A. maculatus, A. potschi, and A. ronnie.
<i>Atractus zebrinus</i> (Jan, 1862) Figs. 18B, 19, 20 <i>Rabdosoma zebrinum<... more <i>Atractus zebrinus</i> (Jan, 1862) Figs. 18B, 19, 20 <i>Rabdosoma zebrinum</i> Jan, 1862; Arch. Zool. Anat. Fis. 2:15. <i>Rhabdosoma maculatus</i> – Günther, 1858; Catalogue of the Colubrine Snakes in the British Museum:421. (part.). <i>Atractus maculatus</i> – Boulenger, 1894; Catalogue of the Snakes in the British Museum 2:306. (part.). <i>Atractus guentheri</i> – Amaral, 1978; Serpentes do Brasil: Iconografia Colorida:114. <i>Atractus</i> cf. <i>guentheri</i> – Lema, 1994; Comun. Mus. Ciênc. Tecnol. PUCRS, Sér. Zool. 7:105. (part.). <i>Atractus maculatus</i> – Moura-Leite, Morato &amp; Bérnils, 1996; Herp. Rev. 27:216. <i>Atractus zebrinus</i> – Fernandes, Freire &amp; Puorto, 2000; Bol. Mus. Nac. Nov. Sér. Zool. 419:4. <b>Holotype:</b> Specimen without defined locality, originally housed at Museo Civico di Storia Naturale di Milano (Jan 1862), believed to have been destroyed during the Second World War (S. Scali pers. comm.). <b>Diagnosis:</b> <i>Atractus zebrinus</i> is distinguished from all congeners by the following combination of characters: (1) 17/17/17 smooth dorsal scale rows; (2) two postoculars; (3) loreal moderate; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) generally seven infralabials, first four contacting chinshields; (7) eight to ten maxillary teeth; (8) generally three gular scale rows; (9) generally three preventrals; (10) 149–170 ventrals in females, 136–153 in males; (11) 15–27 subcaudals in females, 20–32 in males; (12) dorsum creamish red with irregular transverse black blotches with white borders in adults; (13) venter immaculate creamish white; (14) moderate body size, with females reaching 600 mm SVL and males 457 mm SVL; (15) tail short in females (6.8–11.7% SVL), moderate to long (10.0–16.5% SVL) in males; (16) hemipenis slightly or moderate bilobed, semicapitate, and semicalyculate. <b>Comparisons:</b> Among all congeners, <i>A. zebrinus</i> shares 17 dorsal scale rows, seven upper and lower labials, first four infralabiais contacting chinshi [...]
<i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007 Figs. 13C, 15A... more <i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007 Figs. 13C, 15A <i>Atractus ronnie</i> Passos, Fernandes &amp; Borges-Nojosa, 2007; Copeia 2007:789. <i>Atractus ronnie</i> – Loebmann, Ribeiro, Sales &amp; Almeida, 2009; Biotemas 22:170. <b>Holotype:</b> Adult female, MNRJ 14194, from Serra de Baturité (04°10'S, 38°55'W, ca. 800 m), municipality of Pacoti in the state of Ceará, Brazil, collected on 10 April 1998 by D. M. Borges-Nojosa (specimen examined). <b>Paratypes:</b> Twenty-four specimens, all from Serra do Baturité, state of Ceará: municipality of Pacoti (CHUFC 1396, MNRJ 14195), collected on 17 July 1989 by L. W. Lima-Verde; (CHUFC 2646), 18 December 1997 by D. M. Borges-Nojosa; (CHUFC 2648), 11 January 1998 by D. M. Borges-Nojosa; (CHUFC 2658), 09 December 2005 by D. M. Borges-Nojosa, J. C. L. Melo, and M. J. B. Leite, locality of Sítio Olho d`água dos Tangarás; (CHUFC 2481, 3500), 23 February 1989 by D. M. Borges-Nojosa, locality of Monguba; (CHUFC 3502), 03 April 1990 by D. M. Borges-Nojosa, locality of Sítio São José; (MNRJ 14196–97), 16 February 1999 and 10 April 1998, respectively by D. M. Borges-Nojosa, locality of Granja; (CHUFC 2641, 2647), 02 November 1997 by D. M. Borges-Nojosa, locality of Sítio Pau do Alho; (CHUFC 25980), 28 April 2005 by W. C. M. Luz, locality of Sítio Xangrilá; (CHUFC 2652–24), 2005 December by W. C. M. Luz, locality of Cidade Pacoti; (CHUFC 2675–76, 2678), between 17 February and 02 March 2006 by W. C. M. Luz, locality of Cidade Pacoti; (CHUFC 2733), 04 April 1990 by D. M. Borges-Nojosa, district of Santana. Municipality of Baturité: (CHUFC 2578), 11 February 2005 by D. M. Borges-Nojosa, P. Cascon, and J. C. L. Melo, locality of Sítio Escuro. Municipality of Mulungu: (CHUFC 2645), 1991 by D. M. Borges-Nojosa and S. M. Cornélio. Municipality of Guaramiranga: (CHUFC 2649), 15 August 1998 by D. M. Borges-Nojosa, locality of Linha da Serra; (CHUFC 2651), 6 June 2005 by D. M. Borges-Nojosa, P. Cascon, and J. C. L. Melo, locality of Sítio Guaramiranga. <b>Diagno [...]
FIGURE 13. General view of original Sordelli's plate of the Atractus multicinctus. Modified f... more FIGURE 13. General view of original Sordelli's plate of the Atractus multicinctus. Modified from Jan & Sordelli (1865).
FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B)... more FIGURE 9. Sulcate and asulcate sides of the hemipenis of (A) Atractus iridescens (ICN 10902), (B) detailed view of lateral expansion of the hemipenis of Atractus iridesnces, (C) sulcate and asulcate sides of the hemipenis of Atractus typhon sp.nov. (ICN 10901). Scale = 5mm.
FIGURE 10. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) ... more FIGURE 10. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) views of head of the lectotype of Tricheilostoma macrolepis (ZMB 1434).
FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostom... more FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the paratype of Tricheilostoma brevissimum (MCZ 38950).
FIGURE 3. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) v... more FIGURE 3. Dorsal (A) and ventral (B) views of body and dorsal (C), lateral (D), and ventral (E) views of head of the lectotype of Trilepida macrolepis (ZMB 1434). Photos by F. Tillack (ZMB).
(2002) FERNANDES et al. Boletim do Museu nacional. Read by researchers in: 100% Biological Scienc... more (2002) FERNANDES et al. Boletim do Museu nacional. Read by researchers in: 100% Biological Sciences. The taxonomic status of the lowland species of the Liophis cobella group is revised, with the recognition of four species: L. cobella (Linnaeus, 1758), L. breviceps Cope, 1860 ...
The taxonomic status of Atractus sanctaemartae and A. nebularis is revised on the basis of quanti... more The taxonomic status of Atractus sanctaemartae and A. nebularis is revised on the basis of quantitative and qualitative analyses of morphological characters (meristics, morphometrics, colour pattern and hemipenis). Characters used previously for diagnosingAtractus nebularis from A. sanctaemartae are demystified, and the synonymy of these species is proposed. Colour pattern, meristic, morphometrical and scale ornamentation differences originally employed in recognizing both taxa are explained as sexually dimorphic characters, which hold no geographic or phylogenetic basis. Additionally, a new species closely related to Atractus sanctaemartae is described from the Atlantic coast of Colombia. It is distinguished from A. sanctaemartae mainly by having two gular scale rows, slender (62% of body diameter) and acuminate head, snout acuminate in lateral view, dorsum uniformly reddish brown and venter cream with a series of rhomboidal blotches arranged linearly on the lateral portion of the ...
We report on the morphological variation (meristic, morphometric, pholidosis, and color pattern c... more We report on the morphological variation (meristic, morphometric, pholidosis, and color pattern characters) and sexual dimorphism of Trilepida macrolepis. We also took the opportunity to redescribe the holotype of Rena affinis in an effort to evaluate its taxonomic status. Based on the congruence of characters from external morphology, we transfer Stenostoma affine to the genus Trilepida. We also discuss the Andean specimens previously refereed to Trilepida affinis and Trilepida macrolepis and provide an identification key for the genus Trilepida.
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