Italian Botanist 13: 27–44 (2022)
doi: 10.3897/italianbotanist.13.85863
RESEARCH ARTICLE
https://italianbotanist.pensoft.net
Notulae to the Italian alien vascular flora: 13
Gabriele Galasso1, Gianniantonio Domina2, Claudia Angiolini3, Dario Azzaro4,
Gianluigi Bacchetta5, Enrico Banfi1, Davide Barberis6, Giulio Barone2,
Fabrizio Bartolucci7, Alessio Bertolli8, Rossano Bolpagni9, Gianmaria Bonari10,
Luca Bracchetti11, Giacomo Calvia5, Giuliano Campus12, Laura Cancellieri13,
Viviana Cavallaro14, Fabio Conti7, Alba Cuena-Lombraña5, Evelina D’Alessandro15,
Giusy Dal Corso16, Alice Dalla Vecchia9, Antonino De Natale17,
Emanuele Del Guacchio17, Giuseppe Di Gregorio18, Emilio Di Gristina2,
Mario Di Stefano4, Emanuele Fanfarillo3, Alessandro Federici19,
Germano Federici20, Giulio Ferretti21, Francesco Festi8, Tiberio Fiaschi3,
Goffredo Filibeck13, Mauro Fois5, Luca Gariboldi22, Giovanni Gestri23,
Leonardo Gubellini24, Alessandro Guiggi25, Nicole Hofmann26,
Valentina L.A. Laface27, Andrea Lallai5, Valerio Lazzeri28, Angela P. Lecis29,
Michele Lonati6, Fernando Lucchese30, Jacopo Lupoletti31, Samuele Maestri13,
Andrea Mainetti32, Francesca Mantino33, Francesco Mascia5, Rizzieri R. Masin34,
Giacomo Mei27, Marco Merli35, Antonino Messina36, Carmelo M. Musarella27,
Ginevra Nota6, Nicola Olivieri37, Bruno Paura38, Roberto Pellegrini39,
Antonio Pica40, Marco Pittarello6, Lina Podda5, Safiya Praleskouskaya41,
Filippo Prosser8, Giuseppe Ratini42, Simone Ravetto Enri6, Francesco Roma-Marzio43,
Giovanni Salerno30, Alberto Selvaggi44, Adriano Soldano45, Giovanni Spampinato27,
Adriano Stinca46, Federico M. Tardella47, Gianmarco Tavilla4, Valeria Tomaselli33,
Giulia Tomasi8, Luca Tosetto48, Roberto Venanzoni41, Lorenzo Lastrucci49
1 Sezione di Botanica, Museo di Storia Naturale di Milano, Corso Venezia 55, 20121 Milano, Italy
2 Dipartimento di Scienze Agrarie, Alimentari e Forestali (SAAF), Università di Palermo, Viale delle Scien
ze, edificio 4, 90128 Palermo, Italy 3 Dipartimento di Scienze della Vita, Università di Siena, Via P.A.
Mattioli 4, 53100 Siena, Italy 4 Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Università di
Catania, Via A. Longo 19, 95125 Catania, Italy 5 Centro Conservazione Biodiversità (CCB), Dipartimento
di Scienze della Vita e dell’Ambiente (DISVA), Università di Cagliari, Viale Sant’Ignazio da Laconi 13,
09123 Cagliari, Italy 6 Dipartimento di Scienze Agrarie, Forestali e Alimentari (DISAFA), Università di
Torino, Largo P. Braccini 2, 10095 Grugliasco (Torino), Italy 7 Centro Ricerche Floristiche dell’Appennino
(Università di Camerino Parco Nazionale del Gran Sasso e Monti della Laga), San Colombo, 67021 Bari
sciano (L’Aquila), Italy 8 Fondazione Museo Civico di Rovereto, Largo Santa Caterina 41, 38068 Rovereto
(Trento), Italy 9 Dipartimento di Scienze Chimiche, della Vita e della Sostenibilità Ambientale, Università di
Parma, Parco Area delle Scienze 33/a, 43124 Parma, Italy 10 Facoltà di Scienze e Tecnologie, Libera Univer
sità di Bolzano, Piazza Università 5, 39100 Bolzano, Italy 11 Scuola di Bioscienze e Medicina Veterinaria,
Università di Camerino, Via Gentile III da Varano, 62032 Camerino (Macerata), Italy 12 Via G. Rossini
69, 09045 Quartu Sant’Elena (Cagliari), Italy 13 Dipartimento di Scienze Agrarie e Forestali (DAFNE),
Copyright Gabriele Galasso et al.. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC
BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
28
Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
Università della Tuscia, Via San Camillo de Lellis snc, 01100 Viterbo, Italy 14 Museo Orto Botanico, Uni
versità di Bari Aldo Moro, Via E. Orabona 4, 70125 Bari, Italy 15 Piazza Molise 13/f, 86100 Campobasso,
Italy 16 Via G.B. Zenati 15, 37060 Buttapietra (Verona), Italy 17 Dipartimento di Biologia, Università
di Napoli Federico II, Via Vicinale Cupa Cintia 21, 80126 Napoli, Italy 18 Via Monte Bonifato 2, 90146
Palermo, Italy 19 Via Civitanova Marche 25, 25040 Esine (Brescia), Italy 20 Piazza Madonna delle Nevi
17, 24068 Seriate (Bergamo), Italy 21 Dipartimento di Biologia, Università di Firenze, Via G. La Pira 4,
50121 Firenze, Italy 22 Via Ghisolfa 3, 20017 Rho (Milano), Italy 23 Via B. Bonfiglioli 30, 59100 Prato,
Italy 24 Centro Ricerche Floristiche Marche, Provincia di Pesaro e Urbino, Via E. Barsanti 18, 61122 Pesaro
(Pesaro e Urbino), Italy 25 Dipartimento di Scienze della Terra, dell’Ambiente e della Vita (DISTAV), Uni
versità di Genova, Corso Europa 26, 16132 Genova, Italy 26 Dipartimento di Scienze Agrarie, Alimentari
ed Ambientali (D3A), Università Politecnica delle Marche, Via Brecce Bianche 10, 60131 Ancona, Italy
27 Dipartimento di Agraria, Università Mediterranea di Reggio Calabria, Località Feo di Vito snc, 89122
Reggio Calabria, Italy 28 Associazione Amici della Natura, c/o Museo di Storia Naturale di Rosignano Marit
timo, Via E. De Filippo 6, 57016 Rosignano Marittimo (Livorno), Italy 29 Stazione di Uta, Corpo Forestale
e di Vigilanza Ambientale (CFVA), Sesta Strada Ovest, 09068 Uta, fraz. Macchiareddu (Cagliari), Italy
30 Dipartimento di Scienze, Università di Roma Tre, Viale G. Marconi 446, 00146 Roma, Italy 31 Vico
del Sacco 22, 64032 Atri (Teramo), Italy 32 Ufficio Conservazione botanicoforestale, Parco Nazionale Gran
Paradiso, Frazione Valnontey 44, 11012 Cogne (Aosta), Italy 33 Dipartimento di Biologia, Università di
Bari Aldo Moro, Via E. Orabona 4, 70125 Bari, Italy 34 Via Regazzoni Bassa 3, 35036 Montegrotto Terme
(Padova), Italy 35 Via dei Caputéi 7, 38070 Stenico, fraz. Sclemo (Trento), Italy 36 Via G. Giunti 19,
59013 Montemurlo (Prato), Italy 37 Via Maestri del Lavoro 40, 64100 Teramo, Italy 38 Dipartimento
di Agricoltura, Ambiente e Alimenti, Università del Molise, Via F. De Sanctis 1, 86100 Campobasso, Italy
39 Via P. Mascagni 41, 51017 Pescia (Pistoia), Italy 40 Via Strada Storta 11, 66100 Chieti, Italy 41 Di
partimento di Chimica, Biologia e Biotecnologie, Università di Perugia, Via del Giochetto 6, 06122 Perugia,
Italy 42 Via della Lepre 11, 05100 Terni, Italy 43 Sistema Museale di Ateneo, Orto e Museo Botanico, Uni
versità di Pisa, Via L. Ghini 13, 56126 Pisa, Italy 44 Istituto per le Piante da Legno e l’Ambiente (IPLA),
Corso Casale 476, 10132 Torino, Italy 45 Largo Brigata Cagliari 6, 13100 Vercelli, Italy 46 Dipartimento
di Scienze e Tecnologie Ambientali, Biologiche e Farmaceutiche, Università della Campania Luigi Vanvitelli,
Via A. Vivaldi 43, 81100 Caserta, Italy 47 Herbarium Universitatis Camerinensis, Scuola di Bioscienze e
Medicina Veterinaria, Università di Camerino, Via Pontoni 5, 62032 Camerino, Italy 48 Via Pegorina 548,
35040 Casale di Scodosia (Padova), Italy 49 Sistema Museale di Ateneo, Università di Firenze, Via G. La
Pira 4, 50121 Firenze, Italy
Corresponding author: Author (gabriele.galasso@comune.milano.it)
Academic editor: Lorenzo Peruzzi | Received 26 April 2022 | Accepted 3 May 2022 | Published 11 May 2022
Citation: Galasso G, Domina G, Angiolini C, Azzaro D, Bacchetta G, Banfi E, Barberis D, Barone G, Bartolucci
F, Bertolli A, Bolpagni R, Bonari G, Bracchetti L, Calvia G, Campus G, Cancellieri L, Cavallaro V, Conti F, CuenaLombraña A, D’Alessandro E, Dal Corso G, Dalla Vecchia A, De Natale A, Del Guacchio E, Di Gregorio G, Di
Gristina E, Di Stefano M, Fanfarillo E, Federici A, Federici G, Ferretti G, Festi F, Fiaschi T, Filibeck G, Fois M,
Gariboldi L, Gestri G, Gubellini L, Guiggi A, Hofmann N, Laface VLA, Lallai A, Lazzeri V, Lecis AP, Lonati M,
Lucchese F, Lupoletti J, Maestri S, Mainetti A, Mantino F, Mascia F, Masin RR, Mei G, Merli M, Messina A, Musarella
CM, Nota G, Olivieri N, Paura B, Pellegrini R, Pica A, Pittarello M, Podda L, Praleskouskaya S, Prosser F, Ratini G,
Ravetto Enri S, Roma-Marzio F, Salerno G, Selvaggi A, Soldano A, Spampinato G, Stinca A, Tardella FM, Tavilla G,
Tomaselli V, Tomasi G, Tosetto L, Venanzoni R, Lastrucci L (2022) Notulae to the Italian alien vascular flora: 13.
Italian Botanist 13: 27–44. https://doi.org/10.3897/italianbotanist.13.85863
Notulae to the Italian alien vascular flora: 13
29
Abstract
In this contribution, new data concerning the distribution of vascular flora alien to Italy are presented. It
includes new records, confirmations, exclusions, and status changes for Italy or for Italian administrative
regions. Nomenclatural and distribution updates published elsewhere are provided as Suppl. material 1.
Keywords
Alien species, floristic data, Italy, nomenclature
How to contribute
The text for the new records should be submitted electronically to Lorenzo Lastrucci
(lorenzo.lastrucci@unifi.it). The corresponding specimen along with its scan or photograph
has to be sent to FI Herbarium: Museo di Storia Naturale (Botanica), Sistema Museale di
Ateneo, Via G. La Pira 4, 50121 Firenze (Italy). Those texts concerning nomenclatural
novelties (typifications only for accepted names), status changes, exclusions, and
confirmations should be submitted electronically to Gabriele Galasso (gabriele.galasso@
comune.milano.it). Each text should be within 1,000 characters (spaces included).
Floristic records
Acer negundo L. (Sapindaceae)
+ (INV) TOS. – Status change from naturalized to invasive alien for the flora of Toscana.
The presence of Acer negundo is increasingly common in disturbed areas of Toscana.
Numerous individuals colonize urban areas and roadsides (Peruzzi and Bedini 2022+).
From observations made in 2020 in the provinces of Siena and Arezzo, we noticed
that in some areas with a strong disturbance the species grows abundant, sometimes
in dense stands. In Chiusi [(Siena), fraz. Montallese, lungo la strada SP326, WGS84:
43.052790°N, 11.920611°E, margini stradali e ambienti ruderali, 255 m, 17 September
2021, T. Fiaschi, E. Fanfarillo (SIENA)] and other localities (e.g., Monteriggioni (Siena),
WGS84: 43.392968°N, 11.217782°E), we observed abundant, fructifying adult individuals and seedlings. We therefore propose the change of status from naturalized to invasive.
C. Angiolini, E. Fanfarillo, T. Fiaschi
Allium tuberosum Rottler ex Spreng. (Amaryllidaceae)
+ (CAS) LOM: Seriate (Bergamo), Via G. Venezian, rotatoria del ponte sul Fiume
Serio per Via C. Cerioli (CFCE: 0324/1) (WGS84: 45.689694°N, 9.721861°E), scarpata erbosa, 252 m, no exp., 13 November 2021, leg. G. Federici, det. E. Banfi, G.
Galasso (BER-HbFAB No. 107483); ibidem, 15 November 2021, leg. G. Federici, det.
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Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
E. Banfi, G. Galasso (FI, MSNM Nos. 51386, 51387, 51388, 51389). – Casual alien
species new for the flora of Lombardia.
G. Federici, E. Banfi, G. Galasso
Amaryllis belladonna L. (Amaryllidaceae)
+ (CAS) SAR: Carbonia (Sud Sardegna), Domu Beccia (WGS84: 39.223889°N,
8.483611°E), scarpata incolta a bordo strada, 148 m, 16 September 2021, leg. A.
Mammoliti, det. V.L.A. Laface, C.M. Musarella, G. Spampinato (FI). – Casual alien
species new for the flora of Sardegna.
V.L.A. Laface, C.M. Musarella, G. Spampinato
Ambrosia artemisiifolia L. (Asteraceae)
+ (NAT) UMB: Orvieto (Terni), Lago di Corbara (WGS84: 42.726658°N,
12.247858°E), greto sassoso della parte alta delle sponde del Lago di Corbara, con
Xanthium italicum, 135 m, 28 August 2020, R. Venanzoni, S. Praleskouskaya (FI,
PERU barcode PERU-2099). – Naturalized alien species new for the flora of Umbria.
R. Venanzoni, S. Praleskouskaya
Ammannia verticillata (Ard.) Lam. (Lythraceae)
+ (NAT) VEN: Vigasio (Verona), Corte Brigafatta Nuova (WGS84: 45.327162°N,
10.970611°E), risaia, 37 m, 5 September 2021, leg. G. Dal Corso, det. F. Prosser (ROV
No. 80116); ibidem, 7 October 2021, leg. G. Dal Corso, det. F. Prosser (FI). – Naturalized alien species confirmed for the flora of Veneto.
This species was historically widespread in the Venetian ricefields (Argenti et al.
2019), in which today it is almost totally replaced by Ammannia coccinea Rottb. Vigasio is among the localities reported by Goiran (1900) for the province of Verona.
G. Dal Corso, F. Prosser
Bidens aurea (Aiton) Sherff (Asteraceae)
+ (CAS) MAR: Porto Sant’Elpidio (Fermo), Via XX Settembre, nei pressi di un cortile
di una abitazione privata (WGS84: 43.237452°N, 13.756926°E), 21 m, 3 January
2021, A. Pica, J. Lupoletti (FI). – Casual alien species new for the flora of Marche.
The species covered about 10 m2. It probably escaped from flowerbeds along the
main road.
A. Pica, J. Lupoletti
Bidens connata Muhl. ex Willd. (Asteraceae)
+ (NAT) UMB: Perugia (Perugia), fraz. Ponte San Giovanni, sponda destra del Tevere a monte del Nuovo Ponte vecchio (WGS84: 43.09193055°N, 12.45186388°E),
Notulae to the Italian alien vascular flora: 13
31
183 m, 18 October 2020, R. Venanzoni, S. Praleskouskaya (FI, PERU barcode PERU2102). – Naturalized alien species new for the Flora of Umbria.
The species can be found sporadically to frequently along the banks of the river
Tevere in Umbria.
R. Venanzoni, S. Praleskouskaya
Bidens sulphurea (Cav.) Sch.Bip. (Asteraceae)
+ (CAS) TAA: Roncegno Terme (Trento), Cadenzi (WGS84: 46.052056°N,
11.403447°E), margine stradina presso orti, avventizia, 584 m, 18 October 2020, F.
Prosser (FI). – Casual alien species new for the flora of Trentino-Alto Adige.
This species was also observed in San Lorenzo Dorsino (Trento), loc. Glolo (6
September 2021, obs. M. Merli, det. E. Banfi).
F. Prosser, M. Merli, E. Banfi
Cardiospermum halicacabum L. (Sapindaceae)
+ (CAS) LAZ: Roma (Roma), parco pubblico presso Via di Grotta Perfetta (WGS84:
41.828746°N, 12.516325°E), siepe, dove si arrampica su individui di Olea europaea,
Pittosporum tobira e Phyllostachys sp., ca. 254 m, 3 October 2021, N. Olivieri (FI). –
Casual alien species new for the flora of Lazio.
N. Olivieri
Cedrus atlantica (Endl.) G.Manetti ex Carrière (Pinaceae)
+ (CAS) PIE: Almese (Torino), al di sotto della pista tagliafuoco del Musinè
(WGS84: 45.105475°N, 7.442177°E), popolamento misto di latifoglie autoctone
e individui adulti di Cedrus atlantica derivanti da rimboschimento, 350 m, 20 May
2021, D. Barberis, A. Mainetti, G. Nota (FI). – Casual alien species new for the flora
of Piemonte.
Several young individuals were found in a mixed stand with native broadleaf trees
and a few planted Cedrus atlantica trees. The young specimens varied in height from
20 cm to 2 m.
D. Barberis, A. Mainetti, G. Nota
Chamaecyparis lawsoniana (A.Murray) Parl. (Cupressaceae)
+ (CAS) PIE: Germagnano (Torino), loc. Colbeltramo, scarpata della strada che collega Colbeltramo a Castagnole (WGS84: 45.252557°N, 7.440891°E), scarpata stradale,
668 m, 2 April 2021, M. Lonati, M. Pittarello, S. Ravetto Enri (FI). – Casual alien species new for the flora of Piemonte.
A young individual, about 80 cm tall, was found in a shady area under broadleaf trees.
M. Lonati, M. Pittarello, S. Ravetto Enri
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Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
Cotoneaster salicifolius Franch. (Rosaceae)
+ (CAS) MAR: Fermignano (Pesaro e Urbino), loc. Serra Alta (WGS84: 43.659679°N,
12.645607°E), scarpata stradale ai margini di una boscaglia diradata, ca. 340 m, 5
December 2021, N. Hofmann (FI, PESA). – Casual alien species new for the flora
of Marche.
A single individual (2–3 m tall) has been observed.
L. Gubellini, N. Hofmann
Cyperus odoratus L. (Cyperaceae)
+ (INV) ITALIA (UMB): Magione (Perugia), fraz. San Feliciano, Oasi La Valle
(WGS84: 43.106111°N, 12.184722°E), sui resti del canneto degradato (morto!),
vegetazione a dominanza di Cyperus fuscus, 280 m, 18 August 2021, R. Venanzoni,
S. Praleskouskaya (FI, PERU barcode PERU-2091); Tuoro sul Trasimeno (Perugia),
Punta Navaccia (WGS84: 43.189981°N, 12.075511°E), al margine del canneto,
258 m, 18 September 2021, R. Venanzoni, S. Praleskouskaya (PERU barcode PERU2100). – Status change from naturalized to invasive alien for the flora of Italy; invasive
alien species new for the Flora of Umbria.
In Umbria, Cyperus odoratus is found in the strongly nitrophilous muddy beach of
Lake Trasimeno covered by vegetation with Cyperus fuscus L. and Oxybasis chenopiodi
oides (L.) S.Fuentes, Uotila & Borsch. In various sites, it tends to form locally dense
populations of the sublittoral belt.
R. Venanzoni, S. Praleskouskaya
Datura inoxia Mill. (Solanaceae)
– CAM. – Alien species to be excluded from the flora of Campania.
The indication in Galasso et al. (2018) is based on the report for Cilento (“Torre
Ficaiola”, i.e., Spiaggia Ficaiola near Torre Caprioli, Pisciotta municipality, Salerno
province) by De Natale and Strumia (2007), already questioned by Del Guacchio
and La Valva (2017). The relevant specimen (Torre Caprioli, 6 July 1998, Herb. A. De
Natale No. 3671) is to be attributed to Datura wrightii Regel, indeed new to Cilento.
E. Del Guacchio, A. De Natale
Eclipta prostrata (L.) L. (Asteraceae)
+ (NAT) MAR. ‒ Status change from casual to naturalized alien for the flora of Marche.
This species was recorded as casual in the Natural Reserve of Sentina (Conti and
Bracchetti 2016), where it is no longer present due to the erosion of the coast. In
the new locality [Grottammare (Ascoli Piceno), tra Cupra Marittima e Grottammare,
WGS84: 43.008825°N, 13.865765°E, spiaggia, 1 m, 16 October 2018, F. Conti, L.
Bracchetti (APP)], a well-established population can be found.
F. Conti, L. Bracchetti
Notulae to the Italian alien vascular flora: 13
33
Elaeagnus pungens Thunb. (Elaeagnaceae)
+ (CAS) TOS: Pistoia (Pistoia), fraz. Santomoro, sopra il paese presso Case Buchetole
(WGS84: 43.582308°N, 10.584183°E), margine sentiero, ca. 440 m, 12 December
2021, leg. G. Gestri, P. Sandretti, det. G. Gestri. V. Lazzeri (FI). – Casual alien species
new for the flora of Toscana.
An adult plant with flowers and fruits occurs on the edge of a trail, with no current
evidence of spontaneous reproduction.
G. Gestri
Eragrostis pectinacea (Michx.) Nees (Poaceae)
+ (NAT) TOS. – Status change from casual to naturalized alien for the flora of Toscana.
This species is widespread in the lower stretch of the river Tevere [Pieve Santo Stefano (Arezzo), Lago di Montedoglio (WG84: 43.625417°N, 12.073081°E), sponde
melmose originatesi al ritiro delle acque, 384 m, 20 August 2021, R. Venanzoni (FI,
PERU barcode PERU-2113)]. It prefers the large stretches of stony and sandy beaches
that emerge when the water withdraws.
R. Venanzoni, S. Praleskouskaya
Erigeron lilacinus (Sennikov & Kurtto) Sennikov (Asteraceae)
+ (CAS) LOM: Milano (Milano), Parco Porto di Mare (WGS84: 45.431045°N,
9.231650°E), margine erboso, 103 m, no exp., 27 September 2020, G. Galasso, E. Banfi
(MSNM No. 51428); ibidem, Via L. Ornato, lato dei numeri dispari, parcheggi sul lato
della strada (WGS84: 45.520955°N, 9.191965°E), tra gli autobloccanti, 138 m, no exp.,
19 June 2021, G. Galasso (FI). – Casual alien species new for the flora of Lombardia.
G. Galasso, E. Banfi
Euphorbia nutans Lag. (Euphorbiaceae)
+ (NAT) PUG. – Status change from casual to naturalized alien for the flora of Puglia.
The population here reported [Taranto (Taranto), Mar Piccolo, Palude La Vela,
WGS84: 40.476035°N, 17.325862°E, bordo strada, 10 m, 15 October 2021, V. To
maselli, F. Mantino (FI, BI Nos. 42845, 42846, 42847, 42848)] consists of a few thousand individuals, regularly fruiting, widespread along roadsides.
V. Tomaselli, V. Cavallaro, F. Mantino
Fraxinus pennsylvanica Marshall (Oleaceae)
+ (CAS) ITALIA (VEN): Verona (Verona), stazione dei treni di Verona Porta Nuova
(CFCE: 0531/4) (WGS84: 45.428849°N, 10.977357°E), incolto ghiaioso, 64 m, 2
July 2021, leg. F. Prosser, A. Bertolli, G. Tomasi, det. F. Prosser (FI). – Casual alien species new for the flora of Italy (Veneto).
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Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
This species was identified according to Atha and Boom (2017). At least three
50–150 cm tall specimens were observed, born spontaneously from trees growing in
the adjacent Via delle Coste. In some central European countries, F. pennsylvanica is an
invasive species in riparian environments (Verloove 2016).
F. Prosser, G. Tomasi, A. Bertolli
Gamochaeta pensylvanica (Willd.) Cabrera (Asteraceae)
+ (NAT) TAA: Trento (Trento), cortile in Via Vittorio Veneto (CFCE: 9932/2) (WGS84:
46.058487°N, 11.120500°E), aiuola erbosa poco curata, popolazione di oltre 100 esemplari (ca. 25 m2) in fiore, in frutto e sterili, 194 m, 20 October 2021, F. Prosser (FI, ROV
No. 80117). – Naturalized alien species new for the flora of Trentino-Alto Adige.
F. Prosser
Hedera canariensis Willd. (Araliaceae)
+ (NAT) TAA: Bondone (Trento), a W di loc. Olte, a W di Baitoni (CFCE: 0129/3)
(WGS84: 45.806493°N, 10.533911°E), boscaglia, 370 m, 18 October 2011, leg. F.
Prosser, A. Bertolli, det. F. Prosser (FI, ROV No. 61339). – Naturalized alien species new
for the flora of Trentino-Alto Adige.
F. Prosser, A. Bertolli
Hedera hibernica (G.Kirchn.) Bean (Araliaceae)
+ (NAT) VEN: Bardolino (Verona), ca. 1 km a ENE di Bardolino, nella valletta con rivo
a E di Monte Felice (CFCE: 0430/1) (WGS84: 45.554684°N, 10.739976°E), bosco
termofilo, 180 m, 28 February 2004, F. Prosser (FI, ROV No. 44952); Torri del Benaco
(Verona), a E di Punta Pontirola (WGS84: 45.618434°N, 10.701830°E), 240 m, 11
October 2006, leg. F. Prosser, A. Bertolli, det. F. Prosser (ROV No. 50876); Verona
(Verona), fraz. Parona, stazione dei treni (CFCE: 0531/2) (WGS84: 45.477532°N,
10.941611°E), margine di massicciata, inselvatichita, tricomi appressati, in media
con poco meno di 10 raggi, 75 m, 10 September 2020, leg. F. Prosser, A. Bertolli,
G. Tomasi, det. F. Prosser (ROV No. 79310); ibidem, a W di Ca’ di David presso Ca’
Brusà (CFCE: 0631/2) (WGS84: 45.386698°N, 10.962959°E), siepe al margine della
strada, 53 m, 27 October 2020, F. Prosser, G. Tomasi, M. Gecchelin (ROV No. 80038);
Villafranca di Verona (Verona), lungo la linea ferroviaria dismessa a NW di Castel
d’Azzano (CFCE: 0631/2) (WGS84: 45.364584°N, 10.920721°E), inselvatichita nel
cespuglieto, 50 m, 27 October 2020, leg. F. Prosser, G. Tomasi, M. Gecchelin, det.
F. Prosser (ROV No. 79588); Dolcè (Verona), fraz. Peri, stazione dei treni (WGS84:
45.660533°N, 10.901149°E), 129 m, 8 April 2021, A. Bertolli, F. Prosser (ROV No.
80123). – Naturalized alien species new for the flora of Veneto.
Hedera hibernica is quite widespread and was probably, at least partially, confused
with H. canariensis Willd. by Argenti et al. (2019).
F. Prosser, A. Bertolli, F. Festi, G. Tomasi
Notulae to the Italian alien vascular flora: 13
35
Jaborosa integrifolia Lam. (Solanaceae)
+ (NAT) SAR. – Status change from casual to naturalized alien for the flora of
Sardegna.
Since the first report in 2011 (Podda et al. 2012), this species has been observed
naturalized in the countryside of Batteria Boggio (Pula, Cagliari), near its first site
known in loc. Is Loccis Santus (San Giovanni Suergiu, Sud Sardegna) and near Perdaxius and Terraseo (Narcao, Sud Sardegna).
G. Bacchetta, G. Calvia, A. Lallai
Lagarosiphon major (Ridl.) Moss (Hydrocharitaceae)
+ (NAT) EMR: Sissa Trecasali (Parma), porto fluviale di Torricella di Sissa, vicino alla
sponda (WGS84: 44.987640°N, 10.294034°E), profondità 0.5 m, 24 July 2019, R.
Bolpagni, A. Dalla Vecchia (FI barcode FI065222). – Naturalized alien species new for
the flora of Emilia-Romagna.
Lagarosiphon major is one of the most widespread and noxious invasive aquatic
species in Europe and globally (Bolpagni 2021). It is, indeed, included in the European
list of alien species of Union concern [Reg. (EU) no. 1143/2014]. In the river port of
Torricella di Sissa, it forms small monophytic stands.
R. Bolpagni, A. Dalla Vecchia
Miscanthus ×longiberbis (Hack.) Nakai (Poaceae)
+ (CAS) ITALIA (LOM): Iseo (Brescia), a S della rotonda tra Via per Rovato (Via Tangenziale Sud) e Viale Roma (WGS84: 45.649400°N, 10.042614°E), incolto, 191 m,
no exp., 16 October 2021, leg. A. Federici, det. E. Banfi, G. Galasso (FI, MSNM Nos.
51406, 51407). – Casual alien nothospecies new for the flora of Italy (Lombardia).
This is a natural sterile hybrid between Miscanthus sacchariflorus (Maxim.) Benth.
& Hook.f. ex Franch. (China, Korea, Japan) and M. sinensis Andersson (China, SEAsia), no longer cultivated in Italy as an ornamental. It can be distinguished from M.
sinensis by culms up to 2.5(–3.5) m, callus hairs 2–4 times as long as spikelet, spikelets
unawned, and from M. sacchariflorus by pedicels of pedicellate spikelets mostly longer
than 3 mm (Ibaragi et al. 2017).
E. Banfi, G. Galasso, A. Federici
Nymphaea mexicana Zucc. (Nymphaeaceae)
+ (NAT) TOS: Castelnuovo Berardenga (Siena), nei pressi del Mulino di Piermaggiore
sulla strada SP9 di Pievasciata (WGS84: 43.387002°N, 11.405236°E), laghetto artificiale, 272 m, 5 July 2021, leg. G. Bonari, T. Fiaschi, det. C. Angiolini, G. Bonari,
T. Fiaschi (FI, SIENA). – Naturalized alien species new for the flora of Toscana.
This population widely colonizes the surface of an artificial lake. We detected numerous flowering individuals. This species is similar to the sterile hybrid Nymphaea × thiona
36
Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
D.B.Ward, but the two taxa differ, according to Ward (1977), by the dimensions of the
sepals (3–5 cm in N. mexicana vs 5–8 cm in N. × thiona).
C. Angiolini, G. Bonari, T. Fiaschi
Opuntia anacantha Speg. (Cactaceae)
+ (CAS) SAR: Oschiri (Sassari), Diga Coghinas, rupi a ridosso della strada presso la
chiesetta di Santa Barbara (WGS84: 40.788700°N, 9.024265°E), rupi, 205 m, 23 August 2019, G. Calvia (FI). – Casual alien species new for the flora of Sardegna.
In Oschiri the plant produces flowers and fruits, but its reproduction is primarily
vegetative by cladode detachment or by the rooting of sterile fruits.
G. Calvia, A. Guiggi
Opuntia engelmannii Salm-Dyck ex Engelm. (Cactaceae)
+ (NAT) UMB: Amelia (Terni), fraz. Fornole (WGS84: 42.539353°N, 12.459980°E),
incolto con vegetazione a Stipa bromoides e Cymbopogon hirtus, 305 m, S-SE, 22 February 2022, R. Venanzoni (FI, PERU barcode PERU-2109); Montecchio (Terni), strada locale direzione Melezzole-Montecchio, scarpata inferiore della strada (WGS84:
42.668347°N, 12.305811°E), scarpata stradale, 633 m, S, 22 February 2022, R. Venan
zoni (FI, PERU barcode PERU-2108); Spello (Perugia), fraz. Capitan Loreto, strada
SS75bis Foligno-Perugia, prima dell’uscita per Capitan Loreto direzione Perugia, sommità della scarpata destra (WGS84: 43.017289°N, 12.647450°E), scarpata stradale,
210 m, SW, 22 February 2022, leg. R. Venanzoni, det. F. Lucchese, A. Guiggi (FI, PERU
barcode PERU-2110). – Naturalized alien species new for the flora of Umbria.
Populations of this species have been observed for decades.
G. Ratini, A. Guiggi, F. Lucchese, R. Venanzoni
Opuntia pilifera F.A.C.Weber (Cactaceae)
+ (CAS) SIC: Aci Castello (Catania), Scogliera Cannizzaro (WGS84: 37.538144°N,
15.131077°E), scogliera, 20 m, 10 October 2021, leg. G. Spampinato, det. V.L.A. La
face, C.M. Musarella, G. Spampinato (FI, REGGIO). – Casual alien species new for
the flora of Sicilia.
In Sicily, O. pilifera grows along cliffs, on basaltic rocks of volcanic origin, where it
lives with other Opuntia species.
V.L.A. Laface, C.M. Musarella, G. Spampinato
Opuntia spinulifera Salm-Dyck (Cactaceae)
+ (NAT) SIC: Terrasini (Palermo), Baia San Cataldo (WGS84: 38.089721°N,
13.077674°E), sea cliffs, 20 m, 28 April 2021, G. Domina (FI, PAL). – Naturalized
alien species new for the flora of Sicilia.
Notulae to the Italian alien vascular flora: 13
37
A discontinuous area of over 200 m2 near the coast is covered by this species, which reproduces abundantly by seed and vegetatively. In the Favorita Park, in the suburbs of Palermo, some individuals of the same species reproduce by seed and propagate vegetatively.
G. Barone, G. Di Gregorio
Oxalis articulata Savigny (Oxalidaceae)
+ (CAS) MAR. – Alien status indication for the flora of Marche.
Oxalis articulata was indicated for the Marche without invasiveness status by Galasso et al. (2018). However, several individuals, escaped from cultivation, have been
found by us in many localities of the municipalities of Cagli, Cantiano, Piandimeleto,
and Mercatello. According to these observations, O. articulata should be considered
casual for this administrative region.
G. Mei, A. Stinca
Panicum dichotomiflorum Michx. (Poaceae)
+ (NAT) UMB: Baschi (Terni), Lago di Corbara, sponda destra (WGS84: 42.718358°N,
12.258800°E), sponde ciottolose e sabbiose con vegetazione a Xanthium italicum e
Abutilon theophrasti, 134 m, September 2020, leg. R. Venanzoni, S. Praleskouskaya, det.
F.M. Tardella (FI, PERU barcode PERU-2104). – Naturalized alien species new for the
Flora of Umbria.
R. Venanzoni, S. Praleskouskaya, F.M. Tardella
Persicaria filiformis (Thunb.) Nakai (Polygonaceae)
+ (CAS) PIE: Baveno (Verbano-Cusio-Ossola), Via Liberazione (WGS84:
45.911556°N, 8.504567°E), spigolo tra la strada e il marciapiede, 204 m, no exp., 1
October 2021, G. Galasso (FI, MSNM Nos. 51192, 51193); ibidem, leg. A. Selvaggi,
det. G. Galasso (Herb. A. Selvaggi). – Casual alien species new for the flora of Piemonte.
A large clump of individuals of this species was observed.
G. Galasso, L. Gariboldi, A. Selvaggi
Physalis angulata L. (Solanaceae)
+ (NAT) SAR: Villaspeciosa (Sud Sardegna), loc. Cuaddu Moriscu, Lago del Cixerri
(WGS84: 39.275651°N, 8.861169°E), pratelli umidi sulle rive del lago, 39 m, 26 October 2021, A.P. Lecis, F. Mascia, L. Podda (FI, CAG). – Naturalized alien species new
for the flora of Sardegna.
Hundreds of individuals were found sparsely along the shores of the Cixerri dam,
occupying an area of approximately 5,000 m2. We observed a tendency of the species
to colonise also the adjacent areas due to its edible fruits dispersed by animals.
A.P. Lecis, F. Mascia, L. Podda
38
Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
Pinus nigra J.F.Arnold subsp. nigra (Pinaceae)
+ (CAS) SIC: Isnello (Palermo), Madonie, Piano Zucchi, edge of the road SP54
(WGS84: 37.889805°N, 13.999320°E), on a stone wall, 1150 m, 3 June 2021, E. Di
Gristina (FI, PAL). – Casual alien subspecies new for the flora of Sicilia.
Some individuals grow on a vertical stone wall in the Madonie Regional Park, born
from seeds produced by trees growing in a nearby reforested area.
E. Di Gristina, G. Barone
Portulaca grandiflora Hook. (Portulacaceae)
+ (CAS) PUG: Brindisi (Brindisi), Via Taranto tra i numeri civici 3 e 29 (WGS84:
40.633188°N, 17.943848°E), interstizi del marciapiede, 25 m, 23 June 2021, F. Roma
Marzio 1416 (FI, PI No. 044613). – Casual alien species new for the flora of Puglia.
The flowers of Portulaca grandiflora may occur with petals of different colours from
yellow to magenta, and can show a double whorl of petals (Ocampus and Travis Columbus 2012). About ten plants were found, all with yellow petals.
F. Roma-Marzio
Pueraria lobata (Willd.) Ohwi (Fabaceae)
+ (INV) TOS: Scandicci (Firenze), in prossimità dell’ingresso del Centro Don Gnocchi
(WGS84: 43.760264°N, 11.205463°E), area incolta adiacente a un’azienda vivaistica,
in contesto periurbano, 14 September 2021, G. Ferretti (FI); ibidem, 10 November
2021, G. Ferretti (FI). – Invasive alien species new for the flora of Toscana.
Pueraria lobata is a particularly noxious, invasive, densely-packed climbing species, considered among the 100 world’s most invasive alien species (Lowe et al.
2004) and included in the black list of invasive alien species of Union concern [Reg.
(EU) no. 2016/1141]. It has invaded an area of about 200 m2, even climbing on
some trees. This species shows a strong tendency to expand, even when limited by
an urban context.
G. Ferretti
Rubus laciniatus Weston (Rosaceae)
+ (NAT) TOS. – Status change from casual to naturalized alien for the flora of Toscana.
The species was recently recorded as casual alien in Toscana by Canzonieri et al.
(2022). In Agliana (Pistoia) a few fruiting plants of fair size were observed among other
brambles and roses in an extended uncultivated ground periodically mowed only in
the area facing the park (loc. Castelletto, di fronte al centro commerciale di Via E. Berlinguer, WGS84: 43.909620°N, 10.996292°E, ca 45 m, 3 July 2021, A. Messina, FI).
A. Messina, G. Gestri
Notulae to the Italian alien vascular flora: 13
39
Sedum palmeri S.Watson
+ (NAT) LOM. – Status change from casual to naturalized alien for the flora of
Lombardia.
In Milano Sedum palmeri is largely naturalized on the masonry banks of the Naviglio della Martesana, with dozens of individuals for a stretch of at least 1.7 km in
length (WGS84: from 45.514009°N, 9.253945°E to 45.504995°N, 9.237820°E).
G. Galasso, E. Banfi
Setaria adhaerens (Forssk.) Chiov. (Poaceae)
+ (NAT) LOM: Breno (Brescia), loc. Lavarino, a monte della strada (Via Laverino
Inferiore) per Case Bruciate (WGS84: 45.959875°N, 10.315386°E), margine stradale
erboso, ca. 340 m, WNW, 24 October 2020, leg. et det. A. Federici, conf. E. Banfi,
G. Galasso (FI ex Hb. E. Bona No. 071242); ibidem, 9 November 2021, leg. et det. A.
Federici, conf. E. Banfi, G. Galasso (MSNM No. 51401). – Naturalized alien species
new for the flora of Lombardia.
The species was observed also along Via Vigne in Piancogno (Brescia province,
WGS84: 45.929961°N, 10.238172°E, 383 m, SE, 2021, obs. A. Federici).
E. Banfi, A. Federici, G. Galasso
Sicyos angulatus L. (Cucurbitaceae)
+ (NAT) MOL: Oratino (Campobasso), argini del Fiume Biferno in loc. Bivaro
(WGS84: 41.584596°N, 14.559886°E), argine fluviale, 395 m, 25 September 2021,
G. Salerno, E. D’Alessandro, B. Paura (FI, Herb. G. Salerno). – Naturalized alien species
new for the flora of Molise.
G. Salerno, E. D’Alessandro, B. Paura
Solanum sisymbriifolium Lam. (Solanaceae)
+ (NAT) SIC: Pozzallo (Ragusa), Lungomare Pietrenere (WGS84: 36.729444°N,
14.851944°E), aiuole, 3 m, 20 August 2018, A. Soldano 18531 (FI). – Naturalized
alien species new for the flora of Sicilia.
Many individuals were observed along the seafront where the species was missing
in previous years.
A. Soldano
Sporobolus indicus (L.) R.Br. (Poaceae)
+ (NAT) SAR. – Status change from casual to naturalized alien for the flora of
Sardegna.
40
Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
This species was detected in 2016 (Campus and De Pascali 2017) near the
Brotzu Hospital of Cagliari, where currently it has disappeared. In recent years,
we have found it in Nora and Pula (Pula, Cagliari), where it was first introduced
in green lawns and then started to produce viable seeds that are germinating in
neighbouring areas.
M. Fois, A. Cuena-Lombraña, G. Campus
Strigosella africana (L.) Botsch. (Brassicaceae)
+ (CAS) TOS: Montale (Pistoia), sopra Fognano, presso la zona di atterraggio elicotteri
di soccorso (WGS84: 43.572342°N, 11.022752°E), incolto erboso periodicamente
sfalciato, ca. 215 m, 14 April 2019, leg. G. Gestri, det. G. Gestri, V. Lazzeri (FI). –
Casual alien species new for the flora of Toscana.
A few fruiting plants were observed.
G. Gestri, V. Lazzeri
Styphnolobium japonicum (L.) Schott (Fabaceae)
+ (CAS) SIC: Catania (Catania), Via Ipogeo (WGS84: 37.512634°N, 15.079921°E),
bordo strada, 38 m, 26 November 2021, leg. C.M. Musarella, det. V.L.A. Laface,
C.M. Musarella, G. Spampinato (FI, REGGIO). – Casual alien species new for the
flora of Sicilia.
V.L.A. Laface, C.M. Musarella, G. Spampinato
Tagetes minuta L. (Asteraceae)
+ (CAS) TAA: Arco (Trento), tra Massone e Bolognano in loc. Gambor (CFCE:
0032/3) (WGS84: 45.919668°N, 10.908787°E), scarpata incolta, 1 pianta alta
1.5 m, molto ramificata, 150 m, 22 January 2022, leg. M. Merli, det. G. Galasso
(FI, ROV No. 80115, Herb. M Merli). – Casual alien species new for the flora of
Trentino-Alto Adige.
G. Galasso, M. Merli
Tetradium daniellii (Benn.) T.G.Hartley (Rutaceae)
+ (CAS) ITALIA (TAA): Ala (Trento), al termine della stradina presso la sponda destra
del Fiume Adige a NE di Chiesurone (CFCE: 0232/1) (WGS84: 45.786316°N,
11.016831°E), margine stradina nella boscaglia con robinia, 147 m, 10 October 2020,
F. Prosser (FI, ROV No. 79638). – Casual alien species new for the flora of Italy (Trentino-Alto Adige).
This species was identified according to Zhang and Hartley (2008). It is a plant
of melliferous interest, planted near an apiary in the indicated locality. It features
Notulae to the Italian alien vascular flora: 13
41
abundant renewal up to 15 m away. In Europe, it sporadically escapes from cultivation, for example in southwestern Germany (Mazomeit 2016).
F. Prosser
Thinopyrum obtusiflorum (DC.) Banfi (Poaceae)
+ (NAT) LAZ: Cittaducale (Rieti), presso Grotti, strada SR578 “Salto Cicolana”,
presso lo svincolo (WGS84: 42.352123°N, 12.957368°E), incolto ruderale, 405 m,
23 November 2020, G. Filibeck, L. Cancellieri (Herb. L. Cancellieri); ibidem, 22 July
2021, G. Salerno, S. Maestri (FI, Herb. G. Salerno); Castel Madama (Roma), Via San
Quirico (WGS84: 41.976970°N, 12.877654°E), 25 July 2021, G. Salerno (Herb. G.
Salerno). – Naturalized alien species new for the flora of Lazio.
This species was sown in 2016 for “greening” of a highway escarpment at Grotti,
and some individuals now grow also in ruderal meadows nearby. Near Castel Madama,
a large population grows on a highway escarpment and nearby areas. The use of this
species for embankment greening followed by local naturalization is already reported
for N-Italy and Switzerland (Bertolli and Prosser 2014 and references therein; Röthlisberger 2010). A germination test on spikelets collected from the Grotti plants confirmed that the population is able to reproduce.
L. Cancellieri, G. Filibeck, S. Maestri, G. Salerno
Triticum turgidum L. subsp. durum (Desf.) Husn. (Poaceae)
+ (CAS) SIC: Caltagirone (Catania) (WGS84: 37.126671°N, 14.519971°E), lungo
il bordo del sentiero, 329 m, 8 April 2021, D. Azzaro, S. Cambria (FI, CAT); Terrasini (Palermo), Baia San Cataldo (WGS84: 38.088381°N, 13.079131°E), 40 m,
28 April 2021, G. Domina (SAF No. 100039). – Casual alien subspecies new for the
flora of Sicilia.
D. Azzaro, G. Tavilla, M. Di Stefano
Zizania latifolia (Griseb.) Hance ex F.Muell. (Poaceae)
+ (CAS) ITALIA (VEN): Sant’Urbano (Padova), sinistra idrografica del Canale Gorzone, ca. 220 m a E di Ponte Zane (WGS84: 45.143497°N, 11.646712°E), sponda
di stagno, in bosco, 9 m, July 2021, L. Tosetto (FI). – Casual alien species new for the
flora of Italy (Veneto).
This species was identified according to Liu and Phillips (2006). The plant is cultivated in China for food use of the young shoots and rhizomes when infected by the
fungus Ustilago esculenta Henn. (Basidiomycota), which renders the indicated parts edible and inhibits the normal development of the synflorescence. Since the plants found
do not bloom, their presence at the site is almost certainly related to this use.
E. Banfi, L. Tosetto, R.R. Masin, R. Pellegrini
42
Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
Nomenclatural and distribution updates from other literature sources
Nomenclatural, status, distribution updates, and corrections to Galasso et al. (2018)
are provided in Suppl. material 1.
G. Galasso, F. Bartolucci
Acknowledgements
We gratefully acknowledge the Acta Plantarum staff, Nicola M.G. Ardenghi, Maurizio
Bovio, Antonio Giacò, Fabrizio Martini, and Lorenzo Pinzani who provided distribution information.
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Gabriele Galasso et al. / Italian Botanist 13: 27–44 (2022)
Supplementary material 1
Supplementary data
Authors: Gabriele Galasso, Fabrizio Bartolucci
Data type: Species data
Explanation note: 1. Nomenclatural updates; 2. Note updates; 3. Distribution updates; 4. Synonyms, misapplied or included names.
Copyright notice: This dataset is made available under the Open Database License
(http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License
(ODbL) is a license agreement intended to allow users to freely share, modify, and
use this Dataset while maintaining this same freedom for others, provided that the
original source and author(s) are credited.
Link: https://doi.org/10.3897/italianbotanist.13.85863.suppl1