Meaning and Scope of Ecology

Ecology can be defined as the study of organisms, plants and animals in relation to each other and to their non-living environment. The term ecology is derived from the Greek word 'Oekologie'. Literally 'Oekos' means 'home or surroundings' and 'logos' means 'study'. Thus it can be said that ecology is the study of earth's household including plants, animals, micro-organisms and human beings that live together as independent components. SCOPE: Earlier ecology was considered to be a branch of biology that deals with the relation of organisms or group of organisms to their environment. Later on, it was realised that ecology is not only concerned with the organisms, plants and animals but also with the energy flows and material cycles on the lands, in the oceans, in the air and in fresh waters. So keeping with the modern emphasis, ecology is defined as the study of structure and function of nature. The study of ecology starts with the understanding of the interaction of various biological units with their environment for the production of a series of living systems, i.e. Ecosystems. Ecosystems are classified on the basis of type and level of gross energy flow. Now comes the understanding of the flows of energy and exchange of materials between ecosystems. Then movement of materials back and forth between organisms and environment is taken up. This movement of materials is known as Bio-geochemical cycles. The most important object of ecology is the study and maintenance of ecological balance and role of man in the degradation of the environment and to suggest remedies for minimising the effects of pollution, deforestation, population explosion, killing of animals and many other problems. The scope of ecology covers a wide array of interacting levels of organization spanning microlevel (e.g., cells) to planetary scale (e.g., ecosphere) phenomena. Ecosystems, for example, contain populations of individuals that aggregate into distinct ecological communities. It can take thousands of years for ecological processes to bring about the final succession stages of a forest. An ecosystem's area can vary greatly, from tiny to vast. A single tree is of little consequence to the classification of a forest ecosystem, but critically relevant to organisms living in and on it. Several generations of an aphid population can exist over the lifespan of a single leaf. Each of those aphids, in turn, supports diverse bacterial communities. The nature of connections in ecological communities cannot be explained by knowing the details of each species in isolation, because the emergent pattern is neither revealed nor predicted until the ecosystem is studied as an integrated whole. Some ecological principles, however, do exhibit collective properties where

the sum of the components explain the properties of the whole, such as birth rates of a population being equal to the sum of individual births over a designated time frame. The scope of ecology stems into every area of human life; we can see the evidence of ecology or bionomics in society. Human health is a common issue; laws are passed to prevent us from polluting the air and the water. Our automobiles have to meet certain standards to control the amount of emissions being released into the atmosphere. We are encouraged to recycle to stop the landfills from overflowing with pollution that will not degrade. Now in early education, children are taught the scope of ecology; they are taught socialization skills, as well as environmental skills. They are taught the relevance of recycling and composting to the health of their environment. Children who are taught from an early age will grow up to be good stewards of the environment, to preserve it for the next generation, which is the scope of ecology in practice.
Branch The two main branches of Ecology are:

Autecology - the study of the interactions of an individual organism or a single species with the living and nonliving factors of its environment. Autecology is primarily experimental and deals with easily measured variables such as light, humidity, and available nutrients in an effort to understand the needs, life history, and behaviour of the organism or species. Synecology - the ecological study of different natural communities or ecosystems.

Hierarchical ecology
System behaviors must first be arrayed into levels of organization. Behaviors corresponding to higher levels occur at slow rates. Conversely, lower organizational levels exhibit rapid rates. For example, individual tree leaves respond rapidly to momentary changes in light intensity, CO2 concentration, and the like. The growth of the tree responds more slowly and integrates these short-term changes.

The scale of ecological dynamics can operate like a closed system, such as aphids migrating on a single tree, while at the same time remain open with regard to broader scale influences, such as atmosphere or climate. Hence, ecologists classify ecosystems hierarchically by analyzing data collected from finer scale units, such as vegetation associations, climate, and soil types, and integrate this information to identify emergent patterns of uniform organization and processes that operate on local to regional, landscape, and chronological scales. To structure the study of ecology into a conceptually manageable framework, the biological world is organized into a nested hierarchy, ranging in scale from genes, to cells, to tissues, to organs, to organisms, to species, and up to the level of the biosphere. This framework forms a panarchy and exhibits non-linear behaviours; this means that "effect and cause are disproportionate, so that small changes in critical variables, such as the numbers of nitrogen fixers, can lead to disproportionate, perhaps irreversible, changes in the system properties."

Biodiversity
Biodiversity is the variety of life and its processes. It includes the variety of living organisms, the genetic differences among them, the communities and ecosystems in which they occur, and the ecological and evolutionary processes that keep them functioning, yet ever changing and adapting.

Biodiversity (an abbreviation of "biological diversity") describes the diversity of life from genes to ecosystems and spans every level of biological organization. The term has several interpretations, and there are many ways to index, measure, characterize, and represent its complex organization. Biodiversity includes species diversity, ecosystem diversity, genetic diversity and the complex processes operating at and among these respective levels. Biodiversity plays an important role in ecological health as much as it does for human health. Preventing species extinctions is one way to preserve biodiversity, but factors such as genetic diversity and migration routes are equally important and are threatened on global scales. Conservation priorities and management techniques require different approaches and considerations to address the full ecological scope of biodiversity. Populations and species migration, for example, are sensitive indicators of ecosystem services that sustain and contribute natural capital toward the well-being of humanity. An understanding of biodiversity has practical application for ecosystem-based conservation planners as they make ecologically responsible decisions in management recommendations to consultant firms, governments, and industry. The protected areas have been established under the protected area network across the world for conservation of biodiversity.

Habitat
The habitat of a species describes the environment over which a species is known to occur and the type of community that is formed as a result. More specifically, "habitats can be defined as regions in environmental space that are composed of multiple dimensions, each representing a biotic or abiotic environmental variable; that is, any component or characteristic of the environment related directly (e.g. forage biomass and quality) or indirectly (e.g. elevation) to the use of a location by the animal." For example, a habitat might be an aquatic or terrestrial environment that can be further categorized as a montane or alpine ecosystem. Habitat shifts provide important evidence of competition in nature where one population changes relative to the habitats that most other individuals of the species occupy. For example, one population of a species of tropical lizards (Tropidurus hispidus) has a flattened body relative to the main populations that live in open savanna. The population that lives in an isolated rock outcrop hides in crevasses where its flattened body offers a selective advantage. Habitat shifts also occur in the developmental life history of amphibians and in insects that transition from aquatic to terrestrial habitats. Biotope and habitat are sometimes used interchangeably, but the former applies to a community's environment, whereas the latter applies to a species' environment.

Biodiversity of a coral reef. Corals adapt and modify their environment by forming calcium carbonate skeletons. This provides growing conditions for future generations and forms a habitat for many other species.

Niche
Termite mounds with varied heights of chimneys regulate gas exchange, temperature and other environmental parameters that are needed to sustain the internal physiology of the entire colony.

Definitions of the niche date back to 1917, but G. Evelyn Hutchinson made conceptual advances in 1957 by introducing a widely adopted definition: "the set of biotic and abiotic conditions in which a species is able to persist and maintain stable population sizes." The ecological niche is a central concept in the ecology of organisms and is sub-divided into the fundamental and the realized niche. The fundamental niche is the set of environmental conditions under which a species is able to persist. The realized niche is the set of environmental plus ecological conditions under which a species persists. The Hutchinsonian niche is defined more technically as a "Euclidean hyperspace whose dimensions are defined as environmental variables and whose size is a function of the number of values that the environmental values may assume for which an organism has positive fitness." Biogeographical patterns and range distributions are explained or predicted through knowledge of a species' traits and niche requirements. Species have functional traits that are uniquely adapted to the ecological niche. A trait is a measurable property, phenotype, or characteristic of an organism that may influence its survival. Genes play an important role in the interplay of development and environmental expression of traits. Resident species evolve traits that are fitted to the selection pressures of their local environment. This tends to afford them a competitive advantage and discourages similarly adapted species from having an overlapping geographic range. The competitive exclusion principle states that two species cannot coexist indefinitely by living off the same limiting resource; one will always outcompete the other. When similarly

adapted species overlap geographically, closer inspection reveals subtle ecological differences in their habitat or dietary requirements. Some models and empirical studies, however, suggest that disturbances can stabilize the coevolution and shared niche occupancy of similar species inhabiting species-rich communities. The habitat plus the niche is called the ecotope, which is defined as the full range of environmental and biological variables affecting an entire species.
Niche construction

Organisms are subject to environmental pressures, but they also modify their habitats. The regulatory feedback between organisms and their environment can affect conditions from local (e.g., a beaver pond) to global scales, over time and even after death, such as decaying logs or silica skeleton deposits from marine organisms. The process and concept of ecosystem engineering has also been called niche construction. Ecosystem engineers are defined as: "organisms that directly or indirectly modulate the availability of resources to other species, by causing physical state changes in biotic or abiotic materials. In so doing they modify, maintain and create habitats." The ecosystem engineering concept has stimulated a new appreciation for the influence that organisms have on the ecosystem and evolutionary process. The term "niche construction" is more often used in reference to the under-appreciated feedback mechanism of natural selection imparting forces on the abiotic niche. An example of natural selection through ecosystem engineering occurs in the nests of social insects, including ants, bees, wasps, and termites. There is an emergent homeostasis or homeorhesis in the structure of the nest that regulates, maintains and defends the physiology of the entire colony. Termite mounds, for example, maintain a constant internal temperature through the design of air-conditioning chimneys. The structure of the nests themselves are subject to the forces of natural selection. Moreover, a nest can survive over successive generations, so that progeny inherit both genetic material and a legacy niche that was constructed before their time.

Biome
Biomes are larger units of organization that categorize regions of the Earth's ecosystems, mainly according to the structure and composition of vegetation. There are different methods to define the continental boundaries of biomes dominated by different functional types of vegetative communities that are limited in distribution by climate, precipitation, weather and other environmental variables. Biomes include tropical rainforest, temperate broadleaf and mixed forest, temperate deciduous forest, taiga, tundra, hot desert, and polar desert. Other researchers have recently categorized other biomes, such as the human and oceanic microbiomes. To a microbe, the human body is a habitat and a landscape. Microbiomes were discovered largely through advances in molecular genetics, which have revealed a hidden richness of microbial diversity on the planet. The oceanic microbiome plays a significant role in the ecological biogeochemistry of the planet's oceans.

Biosphere
The largest scale of ecological organization is the biosphere: the total sum of ecosystems on the planet. Ecological relationships regulate the flux of energy, nutrients, and climate all the way up to the planetary scale. For example, the dynamic history of the planetary atmosphere's CO2 and O2 composition has been affected by the biogenic flux of gases coming from respiration and photosynthesis, with levels fluctuating over time in relation to the ecology and evolution of plants and animals. Ecological theory has also been used to explain self-emergent regulatory phenomena at the planetary scale: for example, the Gaia hypothesis is an example of holism applied in ecological theory. The Gaia hypothesis states that there is an emergent feedback loop generated by the metabolism of living organisms that maintains the temperature of the Earth and atmospheric conditions within a narrow self-regulating range of tolerance.

Population ecology
Population ecology studies the dynamics of species populations and how these populations interact with the environment. A population consists of individuals of the same species that live, interact and migrate through the same niche and habitat. A primary law of population ecology is the Malthusian growth model which states, "a population will grow (or decline) exponentially as long as the environment experienced by all individuals in the population remains constant." Simplified population models usually start with four variables: death, birth, immigration, and emigration. An example of an introductory population model describes a closed population, such as on an island, where immigration and emigration does not take place. Hypotheses are evaluated with reference to a null hypothesis which states that random processes create the observed data. In these island models, the rate of population change is described by:

where N is the total number of individuals in the population, B is the number of births, D is the number of deaths, b and d are the per capita rates of birth and death respectively, and r is the per capita rate of population change. The formula states that the rate of change in population size (dN/dT) is equal to births minus deaths (B D). Using these modelling techniques, Malthus' population principle of growth was later transformed into a model known as the logistic equation:

where N is the number of individuals measured as biomass density, a is the maximum per-capita rate of change, and K is the carrying capacity of the population. The formula states that the rate

of change in population size (dN/dT) is equal to growth (aN) that is limited by carrying capacity (1 N/K). Population ecology builds upon these introductory models to further understand demographic processes in real study populations. Commonly used types of data include life history, fecundity, and survivorship, and these are analysed using mathematical techniques such as matrix algebra. The information is used for managing wildlife stocks and setting harvest quotas. In cases when the use of null hypotheses is not appropriate, ecologists may adopt different kinds of statistical methods, such as the Akaike information criterion, or use models that can become mathematically complex as "several competing hypotheses are simultaneously confronted with the data."
Metapopulations and migration

The concept of metapopulations was defined in 1969 as "a population of populations which go extinct locally and recolonize." Metapopulation ecology is another statistical approach that is often used in conservation research. Metapopulation models simplify the landscape into patches of varying levels of quality, and metapopulations are linked by the migratory behaviours of organisms. Animal migration is set apart from other kinds of movement because it involves the seasonal departure and return of individuals from a habitat. Migration is also a population-level phenomenon, as with the migration routes followed by plants as they occupied northern postglacial environments. Plant ecologists use pollen records that accumulate and stratify in wetlands to reconstruct the timing of plant migration and dispersal relative to historic and contemporary climates. These migration routes involved an expansion of the range as plant populations expanded from one area to another. There is a larger taxonomy of movement, such as commuting, foraging, territorial behaviour, stasis, and ranging. Dispersal is usually distinguished from migration because it involves the one way permanent movement of individuals from their birth population into another population. In metapopulation terminology, migrating individuals are classed as emigrants (when they leave a region) or immigrants (when they enter a region), and sites are classed either as sources or sinks. A site is a generic term that refers to places where ecologists sample populations, such as ponds or defined sampling areas in a forest. Source patches are productive sites that generate a seasonal supply of juveniles that migrate to other patch locations. Sink patches are unproductive sites that only receive migrants; the population at the site will disappear unless rescued by an adjacent source patch or environmental conditions become more favourable. Metapopulation models examine patch dynamics over time to answer questions about spatial and demographic ecology. The ecology of metapopulations is a dynamic process of extinction and colonization. Small patches of lower quality (i.e., sinks) are maintained or rescued by a seasonal influx of new immigrants. A dynamic metapopulation structure evolves from year to year, where some patches are sinks in dry years and are sources when conditions are more favourable. Ecologists use a mixture of computer models and field studies to explain metapopulation structure.