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Phylum Porifera

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Hyalonema

1.4 Phylum Porifera.


(i) Introduction : Lowest multicellular animals or metazoans without true tissues, i.e.,
at Cellular level of body organization. Familiar as sponges, these animals are well-known
for their ability to absorb and withhold fluids. The word Porifera means pore bearers (Gr.,
porus = pore; ferre = to bear); Their body wall has numerous minute pores, called ostia,
through which a continuous current of outside water is drawn into the body. About 5,000
species are known.
(ii) Brief History : Robert Grant (1825) finally proved that sponges are animals, and
coined the name Porifera for these. Schulze (1878), Butschli (1884), Sollas (1884) and
Delage (1898) separated sponges from other metazoans on basis of embryological studies,
and suggested a separate group, Parazoa for these.
(iii) Salient features : Phylum porifera has the following salient features :
(1) All the sponges are Aquatic, Sedentary, Asymmetrical or Radially, First
multicellular organisms and have cellular grade of organization.
(2) They are diploblastic. Ectoderm is formed by pinachocyte and endoderm is formed
by choanocyte. Both layers are called pinachoderm and choandoderm.
(3) The body is perforated by numerous minute pores called ostia.
(4) The ostia open into a large cavity called spongocoel.
(5) The spongocoel opens to the outside by a large opening called osculum.
(6) The sponges possess an endoskeleton in the form of calcareous spicules.
(7) Excretion and respiration occur by diffusion.
(8) They have greater power of regeneration.
(9) Reproduction takes place by asexual or sexual methods.
(10) Development is indirect or direct. The common larval are parenchymula,
amphiblastula, etc.
(a) Class 1. Calcarea
(1) Skeleton is formed of Calcareous spicules.
(2) Radially symmetrical.
(3) Choanocyte cells are large and conspicuous
Examples : Clathrina, Leucosolenia, Sycon, etc.,
Euplectela is the sponge which is given as a Gift in Japan.
Leucosolenia is a smallest sponge.
Ectorderm is formed by pinachocyte and endoderm is
formed by choanocyte. Both layer is called pinachoderm and
choanoderm.
(b) Class 2. Hexactinellida
(1) Skeleton is formed of six rayed triaxon, silicious spicules,
(2) Canal system is branched or unbranched.
(3) Radially symmetrical.
(4) These are also known as glass sponges.
Examples : Pheronema, Hyalonema, etc.,
Hylonema is a Glass rope sponge.
(c) Class 3. Demospongia
(1) Skeleton either absent or present. When present it is either formed of spongin fibres
or combination of spongin fibres and silicious spicules.
(2) The silicious spicules when present are never six rayed
(3) The canal system is complicated Rhagon type
(4) These sponges are of great economic importance
Euspongia
Sycon
Examples : Cliona, Spongilla, Chalina, Euspongia, Hippospongia, Oscarella, etc.,
Spongilla is a fresh water sponge.

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