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Special Senses (EAR) : Sumera Afzal

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SPECIAL SENSES

(EAR)

SUMERA AFZAL
SENIOR LECTURER
ZCRS
FUNCTIONS OF EAR

 Hearing
 Balance & equilibrium
STRUCTURE OF EAR
 Anatomically divided into 3 parts:

 External ear

 Middle ear

 Inner ear
EXTERNAL EAR

COMPONENTS:
 Pinna / Auricle

 External auditory meatus / Auditory canal

 Tympanic membrane / Ear drum


FUNCTIONS:
 Protection
 Amplification
 Receives sound waves
 Pass the sound waves to external auditory meatus
 From external auditory meatus to tympanic
membrane
 Tympanic membrane connects external ear to middle
ear
MIDDLE EAR
 An air filled cavity in temporal bone
COMPONENTS:
 Eustachian (auditory) tube
 Auditory Ossicles
 Malleus
 Incus
 Stapes
Eustachian Tube
 The middle ear is air filled space
 It consist of both bone and hyaline cartilage
 This runs from the middle ear to naso-pharnyx behind
the nose
 It is closed at the pharyngeal end
 During swallowing, chewing and yawning it opens
 It helps maintaining the equal air pressure on both
sides of the ear drum
 If pressure is disturbed, it hinders clear and normal
hearing
 Manubrium (handle of malleus) is attached to
back of tympanic membrane
 Head is attached to wall of middle ear
 Short process is attached to incus.
 Hence, when malleus moves, incus also moves
with it
 Processes of incus are attached to head of stapes
 Thus, Incus articulates with stapes.
 Faceplate/ footplate of stapes is attached to
opening of oval window lying against the
membranous labyrinth of cochlea by Annular
ligaments
 2 Skeletal muscles support function of hearing:
 Tensor tympani muscles
 Stapedius
 Both of these muscles are located in middle ear
 Tensor tympani muscles are located at the point of attachment
of malleus manubrium with tympanic membrane
 Malleus manubrium is attached to center of tympanic
membrane
 These muscles keep the tympanic membrane tensed
 When the tensor tympani contracts, the malleus is pulled
medially, stiffening the tympanic membrane.
 This increased stiffness reduces the amount of movement
possible
 Stapedius muscles contraction keep the footplate of stapes out
of the oval window
 Contraction of the stapedius pulls the stapes, reducing
movement of the stapes at the oval window.
 Ossicles are suspended by ligaments
 Making Malleus, Incus & Stapes to act as ‘One single
lever’.
 Whereas, articulation of Incus with Stapes cause the
stapes to:

 Push forward on oval window & on cochlear fluid


on the other side of window
Condition: When tympanic membrane moves
inward

 Pull backward on the fluid


Condition: Every time when malleus moves
outward
IMPEDANCE MATCHING
 Impedance: Medium's resistance to movement

 Function of middle ear: Match relatively low-impedance


airborne sounds to the higher-impedance fluid of the inner
ear
 Auditory transduction is the transformation of sound
pressure into electrical energy

 External & middle ear are air filled, where as the inner ear
is fluid filled.

 Thus for auditory transduction, sound waves traveling


through air must be converted into pressure waves in the
fluid of inner ear.
 The tympanic membrane and the chain of ossicles serve
as an impedance-matching device.

 The ear must detect sound waves traveling in air, but the
neural transduction mechanism depends on movement in
the fluid-filled cochlea, where acoustic impedance is
much higher than that of air.

 Therefore, without a special device for impedance


matching, most sound reaching the ear would simply be
reflected, as are voices from shore when a person is
swimming under water.
 Impedance matching in the ear depends on:

(1) The ratio of the surface area of the large tympanic


membrane to that of the smaller oval window

(2) The mechanical advantage of the lever system formed


by the ossicles.

• This impedance matching is sufficient to increase the


efficiency of energy transfer by nearly 30 dB in the range
of hearing from 300 to 3500Hz i.e. usual range of
frequencies in human speech with sound intensity of
about 65 dB .
 Amplitude of movement of stapes faceplate is only 3/4th
as much as amplitude of the handle of malleus

 Therefore, the ossicular lever system does not increase


the movement distance of the stapes, as is commonly
believed.

 Instead, the system actually reduces the distance but


increases the force of movement about 1.3 times

 What that means is that the stapes is displaced much less


than Tympanic membrane
 Surface area of tympanic membrane is 55 mm2

 Surface area of stapes is 3.2 mm2

 This creates a difference of 17 folds

 Thus, area of the tympanic membrane is larger than that of


the stapes foot plate in the cochlea.

 This 17 fold difference in area times (multiplied by) the 1.3


fold ratio of the lever system causes about 22 times as
much total force to be exerted on the fluid of the cochlea as
is exerted by the sound waves against the tympanic
membrane
Therefore, 2 steps in Impedance matching are:

1. Forces collected over the ear drum are now concentrated


over a smaller area, thus increasing the pressure at the
oval window

 Total sound force is increased by 22 times.


2. Lever action of middle ear bones

 Arm of incus is shorter than arm of malleus

 Tympanic membrane act as fulcrum

 Tensor tympani muscle act as effort arm

 Malleus, incus & stapes act as weight arm

“If the effort arm is far, force is increased ”


 In the absence of the ossicular system and tympanic
membrane, sound waves can still travel directly through
the air of the middle ear and enter the cochlea at the oval
window.

 However, the sensitivity for hearing is then 15 to 20


decibels less than for ossicular transmission equivalent to
a decrease from a medium to a barely perceptible voice
level.
ATTENUATION REFLEX
 It is characterized by involuntary contraction of
tensor tympani and stapedius muscles, in response to
a loud noise.
 Has a latent period of 40 to 80 millisecond.
 When both the muscles contract, manubrium of
malleus moves inward and stapes is pulled outward.
 These two actions result in stiffness of auditory
ossicles, so that the transmission of sound is
decreased.
SIGNIFICANCE OF ATTENUATION
REFLEX
i. Tympanic reflex protects the tympanic membrane
from being ruptured by loud sound.

ii. It also prevents fixation of footplate of stapes, against


oval window, during exposure to loud sound.

iii. It helps to protect the cochlea from damaging effects


of loud sounds. Contraction of tensor tympani and
stapedius during exposure to loud sound develops
stiffness of the auditory ossicles so that, the
transmission of sound into cochlea is decreased
INNER EAR
 Also called Labyrinth (Maze)
 Made up of 2 parts one within the other:
 Bony labyrinth
 Membranous labyrinth
 Bony labyrinth contains Perilymph
 Membranous labyrinth is present inside these channels
of bony labyrinth
 Membranous labyrinth contains
Endolymph.
COMPONENTS OF LABYRINTH:

1. Cochlea
2. Semicircular canals
3. Otolith organs

Otolith organs
STRUCTURE OF COCHLEA
 Cochlea is a coiled tube, 35 mm long
 Consists of 3 tubes side by side in each turn:

1. Scala vestibuli
2. Scala media
3. Scala tympani
REISSNER’S PERILYMPH
MEMBRANE
OVAL WINDOW

SCALA VESTIBULI

SCALA MEDIA

SCALA TYMPANI

ROUND WINDOW
BASILAR
ENDOLYMPH
MEMBRANE
Sound vibrations move from faceplate of
stapes into oval window
From oval window into Scala Vestibuli
Stapes, thus moves inward and outward
 Inward movement cause the fluid to move
in Scala vestibuli and Scala media
 Outward movement cause the fluid to
move backward
 Reissner’s membrane do not obstruct sound
movement from Scala vestibuli to Scala media

 On the surface of basilar membrane lies Organ of


Corti
ORGAN OF CORTI
 Organ of corti rests upon the basilar membrane.
 It extends throughout the cochlear duct, except for a short
distance on either end.
 Roof of the organ of Corti is formed by gelatinous
tectorial membrane.
 Organ of Corti is made up of sensory elements called hair
cells and various supporting cells.
 All the cells of organ of Corti are arranged in order from
center towards the periphery of the cochlea.
COMPONENTS OF ORGAN OF CORTI:

 Tectorial membrane
 Hair cells (Inner & Outer)  RECEPTORS
 Pillar cells (Rods of Corti) SUPPORTING
CELLS
 Tunnels of Corti
 Reticular lamina
 Nerve fibers
NERVE INNERVATION OF COCHLEA:
 Afferent neuron that innervates hair cells form the
auditory division of 8th CN  Vestibulocochlear
Nerve
 7th CN Facial Motor Nerve: Tensor tympani
muscles
 5th CN Trigeminal Motor nerve: Stapedius
muscles
AUDITORY PATHWAY
Afferent fibers from 8th cranial nerve

Ends in Dorsal & Ventral Cochlear nuclei located in upper part of


Medulla

Fibers synapse & 2nd order neurons pass mainly to opposite side
of brain stem to terminate in Superior Olivary nucleus

Pass through lateral leminiscus nucleus into inferior colliculus


and synapse again

Fibers now pass to medial geniculate nucleus to synapse again

Fibers finally enter in Primary Auditory cortex located in


Superior Gyrus of temporal lobe
H
D
VESTIBULAR SYSTEM
 Two components are:

 3 Semicircular canals (Anterior, Posterior &


lateral)

 2 Otolith organs (Utricle & Saccule)

 Together they provide constant feedback to the


cerebellum about head movement
 The Otolith organs are sensitive to changes in the
position of the head with respect to gravity (linear
acceleration in vertical or horizontal plane)
e.g. Walking in a row or moving in lift

 Send messages to the brain about body movement in


a straight line (backwards/forwards or
upwards/downwards)
 Also about where the head is in relation to gravity
e.g. Tilting, leaning or lying down
 Sensory epithelium is called Macula
 Contains Supporting cells & Hair cells
 Surrounded by Otolithic membrane
 Membrane contains crystals of Calcium Carbonate-
Otoliths/ Ear dust/ Otoconia
S
s
S
s
 Processes of hair cells are in the Macula

 Afferent nerve fiber here is Vestibular division of C-8


nerve

 These crystals are displaced during head movements


to stimulate tiny hairs, which transmit the message
via the vestibular nerve to the cerebellum.
 The Semicircular canals
are sensitive to angular
accelerations.
e.g. Nodding, rotating

 Receptor structure is
Crista ampularis
 Located on Ampulla
(expanded end of each
membranous canals)
 Crista contains hair cells & supporting
(sustentacular) cells surrounded by Cupula
(Gelatinous partition)

 Processes of hair cells are embedded in ampulla

 Afferent nerve fiber here is Vestibular division of


C-8 nerve
 Canals are stimulated by the weight of the fluid
they contain & the pressure it exerts varying with
the head position.

 Movement of fluid inside the canals caused by


head movement stimulates tiny hairs that send
messages via the vestibular nerve to the
cerebellum.
BALANCE MAINTAINANCE

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