<i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp... more <i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp. nov. (Figs 1, 2) <i>Similis</i> A.austrokhorasanico <i>Podlech sed differt stipulis 16-23 mm longis (nec 10-12 mm), foliolis 7-8 (nec 5-7)-jugis,pedunculo 12-18 (nec 6-11) cm longo, inflorescentiis 3.5-5 (7) cm longis, atque ovario breve stipitato, nullo modo sessili.</i> TYPUS. — <b>Iran</b>. Khorassan province, NW Darrehgaz, Tandureh National Park, Shekarab, western altitudes of Shekarab station, 2200 m, 37°22'45.8''N &amp; 58°42'07.7''E, 27. VI.2004, <i>Memariani &amp; Zangooei 44249</i> (holo-, FUMH!; iso- FAR!, T!). DISTRIBUTION. — <i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp. nov. is endemic to Iran and known only from the type locality, in the Khorassan province, E. Iran (Fig. 3). Th e species is thriving in mountains of Shekarab, Tandureh National Park, at an altitude <i>c.</i> 2200 m, a mountainous area covered with a mixture of gravel and clay, in the Irano-Turanian floristic region (Takhtajan 1986). Th e flowering and fruiting material was collected in June. ETYMOLOGY. — Th e specific epithet is derived from the type locality, East Iran. DESCRIPTION Plant caespitose, spiny, <i>c.</i> 18-32 cm tall, covered with solely white, basifixed hairs. Caudex <i>c.</i> 5-8 mm in diameter. Stems 7-15 cm long, the lower parts ligneous, branched, densely covered with blackish remnants of old leaves. Stipules membranaceous, yellowish, ± hyaline towards the tips, 16-23 mm, adnate to the petiole for 11-18 mm, connate behind the stem for 4-6 mm, the free tips acuminate, many-nerved, glabrous or ciliate at the margins with ascending hairs up to 1mm. Leaves 4.5-7.5 cm; petiole 1-2 cm, like the rachis densely covered with appressed to subappressed-ascending hairs ca. 0.5 mm, spiny. Leaflets in 7-8 pairs, narrowly elliptic, 10-16 × 1.5-3 mm, acuminate, pungent, with a cusp <i>c.</i> 1.5-2 mm, on the both sides densely covered with appressed to subappressed-ascending hairs. Peduncles 12-18 cm, longer than t [...]
The classification of the genus Salix (Salicaceae) is troubled by the intra-specific flexibility ... more The classification of the genus Salix (Salicaceae) is troubled by the intra-specific flexibility and diversity of the morphological features. The aim was to prepare a comparative description of petiole anatomy of 23 species of Salix, in addition to two species of Populus and Bivinia jalbertii, as well as to determine the taxonomic importance of petiole features. In order to do so, 10 quantitative and 10 qualitative petiole anatomical features were studied. Moreover, a Bayesian analysis was executed on the relevant ITS sequences obtained from GenBank, included 28 specimens from 17 species. The anatomical features were then mapped on the resulted cladogram and the ancestral states of some prominent petiole anatomical features were reconstructed. Based on the results, no stomata were observed in the petiole's epidermis of any of the species studied; therefore, it could be assumed that the petiole in the studied species is dependent to the leaf blade for respiration and gas exchange...
A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Ki... more A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Kickxia iranica is morphologically similar to K. commutata, a species native to Mediterranean region, however it can be distinguished from the latter by leaf shape, color of corolla, habit, stem branching and length of leaf, petiole, pedicel and corolla. Notes on its distribution, ecology and phenology are also presented.
Silene ghahremaninejadii (Caryophyllaceae) is described as a new species to science in this artic... more Silene ghahremaninejadii (Caryophyllaceae) is described as a new species to science in this article based on critical differences in certain characters of both vegetative and reproductive organs, compared with its closely related taxa, i.e. S. marschallii and S. ruprechtii. The new species has compact caudex, glabrous leaves and pedicles, cylindrical calyx without indumentums on its inner surface, deeply bifid petals without coronal scales and glabrous filaments. In addition, a hypothesis about the speciation process of the species is proposed.
Large and diverse genera, such as Silene need more reliable morphological traits for the credible... more Large and diverse genera, such as Silene need more reliable morphological traits for the credible identification and delimitation of the species. Despite the fact that the type of indumentum among the species of Silene had been addressed in most available revisions, monographs and floristic studies, the trichome-based features and their adaptational importance have not been investigated explicitly. In the present study, the trichomes of annual species of Silene in Iran are studied. Beside other floral traits and vegetative features, the presence, distribution and mixture of the glandular and eglandular trichomes on stems, leaves, inflorescence axes, pedicels, anthophores, inner and outer surfaces of calyces, petals and styles are proved to be of diagnostic importance in Silene. Indumentum of the inner calyx among the studied species is investigated here for the first time. An identification key is performed mainly based on indumentum features. In addition, a putative trichome-based ...
The genus Bupleurum L., with about 185-195 species, is the second largest genus of the family Api... more The genus Bupleurum L., with about 185-195 species, is the second largest genus of the family Apiaceae Lindl. It includes 35 species in the Iranian plateau, 15 of which are thrived in Iran. Three species out of these, i.e., B. wolffianum, B. flexile and B. ghahremanii, are endemic to Iran. In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum, for which specimens were not available, were studied for the first time. Some of the most taxonomically important fruit characteristics were found to be: mericarp shape in transverse section, characteristics of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important features for each species were described and an identification key was also provided. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and ot...
Having been inhabited for nearly 12,000 years (Matthews et al., 2010) and hosted agricultural soc... more Having been inhabited for nearly 12,000 years (Matthews et al., 2010) and hosted agricultural societies for more than 11 millennia, the plateau of Iran has been enriched in domesticated cultivars, spices and medicinal herbs (Riehl et al., 2013). Experimental methods, such as trial and error, provided human populations with some pieces of data on the usage of their local plants. This data, which has been obtained with much sweat and blood, has passed through generations and seeded the early stages of regional cultures and civilizations. The flora of Iran consists of nearly 8000 wild plant species (Ghahremaninejad, 2015), growing in different environments from wet jungles to arid deserts and from low plains to high mountains of the country. This extensive variety of habitats has made up the stunning biodiversity of the country, which results in the possibility of natural growth or cultivation of various medical and aromatic plants. Identification of medicinal and aromatic plants of Ir...
A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Ki... more A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Kickxia iranica is morphologically similar to K. commutata, a species native to Mediterranean region, however it can be distinguished from the latter by leaf shape, color of corolla, habit, stem branching and length of leaf, petiole, pedicel and corolla. Notes on its distribution, ecology and phenology are also presented.
The genus Bupleurum L, with about 185–195 species, is the second largest genus of the family Apia... more The genus Bupleurum L, with about 185–195 species, is the second largest genus of the family Apiaceae Lindl. It has 35 species in the Iranian plateau, of which 15 are thrived in Iran and three of them, B. wolffianum, B. flexile and B. ghahremanii, are endemic species of to Iran.
In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum due to lack of available specimens, were studied for the first time. Some of the most taxonomically important fruit characteristics are as follows: mericarp shape in transverse section, characteristic of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important characters for each species are described and an identification key is provided to portray the similarities and relationships between the species. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and other members of the genus. Absence of vittae in Sect. perfoliata may be correlated with broad leaves and dilated, rounded and perfoliate leaf bases. Furthermore, two distinct subgroups were shown due to different number of vallecular vittae in each furrow, which may be applied as a basic classification factor for further investigations on the genus Bupleurum.
<i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp... more <i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp. nov. (Figs 1, 2) <i>Similis</i> A.austrokhorasanico <i>Podlech sed differt stipulis 16-23 mm longis (nec 10-12 mm), foliolis 7-8 (nec 5-7)-jugis,pedunculo 12-18 (nec 6-11) cm longo, inflorescentiis 3.5-5 (7) cm longis, atque ovario breve stipitato, nullo modo sessili.</i> TYPUS. — <b>Iran</b>. Khorassan province, NW Darrehgaz, Tandureh National Park, Shekarab, western altitudes of Shekarab station, 2200 m, 37°22'45.8''N &amp; 58°42'07.7''E, 27. VI.2004, <i>Memariani &amp; Zangooei 44249</i> (holo-, FUMH!; iso- FAR!, T!). DISTRIBUTION. — <i>Astragalus orientopersicus</i> F.Ghahrem., Joharchi, Fereid. &amp; Hoseini, sp. nov. is endemic to Iran and known only from the type locality, in the Khorassan province, E. Iran (Fig. 3). Th e species is thriving in mountains of Shekarab, Tandureh National Park, at an altitude <i>c.</i> 2200 m, a mountainous area covered with a mixture of gravel and clay, in the Irano-Turanian floristic region (Takhtajan 1986). Th e flowering and fruiting material was collected in June. ETYMOLOGY. — Th e specific epithet is derived from the type locality, East Iran. DESCRIPTION Plant caespitose, spiny, <i>c.</i> 18-32 cm tall, covered with solely white, basifixed hairs. Caudex <i>c.</i> 5-8 mm in diameter. Stems 7-15 cm long, the lower parts ligneous, branched, densely covered with blackish remnants of old leaves. Stipules membranaceous, yellowish, ± hyaline towards the tips, 16-23 mm, adnate to the petiole for 11-18 mm, connate behind the stem for 4-6 mm, the free tips acuminate, many-nerved, glabrous or ciliate at the margins with ascending hairs up to 1mm. Leaves 4.5-7.5 cm; petiole 1-2 cm, like the rachis densely covered with appressed to subappressed-ascending hairs ca. 0.5 mm, spiny. Leaflets in 7-8 pairs, narrowly elliptic, 10-16 × 1.5-3 mm, acuminate, pungent, with a cusp <i>c.</i> 1.5-2 mm, on the both sides densely covered with appressed to subappressed-ascending hairs. Peduncles 12-18 cm, longer than t [...]
The classification of the genus Salix (Salicaceae) is troubled by the intra-specific flexibility ... more The classification of the genus Salix (Salicaceae) is troubled by the intra-specific flexibility and diversity of the morphological features. The aim was to prepare a comparative description of petiole anatomy of 23 species of Salix, in addition to two species of Populus and Bivinia jalbertii, as well as to determine the taxonomic importance of petiole features. In order to do so, 10 quantitative and 10 qualitative petiole anatomical features were studied. Moreover, a Bayesian analysis was executed on the relevant ITS sequences obtained from GenBank, included 28 specimens from 17 species. The anatomical features were then mapped on the resulted cladogram and the ancestral states of some prominent petiole anatomical features were reconstructed. Based on the results, no stomata were observed in the petiole's epidermis of any of the species studied; therefore, it could be assumed that the petiole in the studied species is dependent to the leaf blade for respiration and gas exchange...
A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Ki... more A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Kickxia iranica is morphologically similar to K. commutata, a species native to Mediterranean region, however it can be distinguished from the latter by leaf shape, color of corolla, habit, stem branching and length of leaf, petiole, pedicel and corolla. Notes on its distribution, ecology and phenology are also presented.
Silene ghahremaninejadii (Caryophyllaceae) is described as a new species to science in this artic... more Silene ghahremaninejadii (Caryophyllaceae) is described as a new species to science in this article based on critical differences in certain characters of both vegetative and reproductive organs, compared with its closely related taxa, i.e. S. marschallii and S. ruprechtii. The new species has compact caudex, glabrous leaves and pedicles, cylindrical calyx without indumentums on its inner surface, deeply bifid petals without coronal scales and glabrous filaments. In addition, a hypothesis about the speciation process of the species is proposed.
Large and diverse genera, such as Silene need more reliable morphological traits for the credible... more Large and diverse genera, such as Silene need more reliable morphological traits for the credible identification and delimitation of the species. Despite the fact that the type of indumentum among the species of Silene had been addressed in most available revisions, monographs and floristic studies, the trichome-based features and their adaptational importance have not been investigated explicitly. In the present study, the trichomes of annual species of Silene in Iran are studied. Beside other floral traits and vegetative features, the presence, distribution and mixture of the glandular and eglandular trichomes on stems, leaves, inflorescence axes, pedicels, anthophores, inner and outer surfaces of calyces, petals and styles are proved to be of diagnostic importance in Silene. Indumentum of the inner calyx among the studied species is investigated here for the first time. An identification key is performed mainly based on indumentum features. In addition, a putative trichome-based ...
The genus Bupleurum L., with about 185-195 species, is the second largest genus of the family Api... more The genus Bupleurum L., with about 185-195 species, is the second largest genus of the family Apiaceae Lindl. It includes 35 species in the Iranian plateau, 15 of which are thrived in Iran. Three species out of these, i.e., B. wolffianum, B. flexile and B. ghahremanii, are endemic to Iran. In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum, for which specimens were not available, were studied for the first time. Some of the most taxonomically important fruit characteristics were found to be: mericarp shape in transverse section, characteristics of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important features for each species were described and an identification key was also provided. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and ot...
Having been inhabited for nearly 12,000 years (Matthews et al., 2010) and hosted agricultural soc... more Having been inhabited for nearly 12,000 years (Matthews et al., 2010) and hosted agricultural societies for more than 11 millennia, the plateau of Iran has been enriched in domesticated cultivars, spices and medicinal herbs (Riehl et al., 2013). Experimental methods, such as trial and error, provided human populations with some pieces of data on the usage of their local plants. This data, which has been obtained with much sweat and blood, has passed through generations and seeded the early stages of regional cultures and civilizations. The flora of Iran consists of nearly 8000 wild plant species (Ghahremaninejad, 2015), growing in different environments from wet jungles to arid deserts and from low plains to high mountains of the country. This extensive variety of habitats has made up the stunning biodiversity of the country, which results in the possibility of natural growth or cultivation of various medical and aromatic plants. Identification of medicinal and aromatic plants of Ir...
A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Ki... more A new species from Iran is described and illustrated here as Kickxia iranica (Plantaginaceae). Kickxia iranica is morphologically similar to K. commutata, a species native to Mediterranean region, however it can be distinguished from the latter by leaf shape, color of corolla, habit, stem branching and length of leaf, petiole, pedicel and corolla. Notes on its distribution, ecology and phenology are also presented.
The genus Bupleurum L, with about 185–195 species, is the second largest genus of the family Apia... more The genus Bupleurum L, with about 185–195 species, is the second largest genus of the family Apiaceae Lindl. It has 35 species in the Iranian plateau, of which 15 are thrived in Iran and three of them, B. wolffianum, B. flexile and B. ghahremanii, are endemic species of to Iran.
In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum due to lack of available specimens, were studied for the first time. Some of the most taxonomically important fruit characteristics are as follows: mericarp shape in transverse section, characteristic of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important characters for each species are described and an identification key is provided to portray the similarities and relationships between the species. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and other members of the genus. Absence of vittae in Sect. perfoliata may be correlated with broad leaves and dilated, rounded and perfoliate leaf bases. Furthermore, two distinct subgroups were shown due to different number of vallecular vittae in each furrow, which may be applied as a basic classification factor for further investigations on the genus Bupleurum.
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In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum due to lack of available specimens, were studied for the first time. Some of the most taxonomically important fruit characteristics are as follows: mericarp shape in transverse section, characteristic of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important characters for each species are described and an identification key is provided to portray the similarities and relationships between the species. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and other members of the genus. Absence of vittae in Sect. perfoliata may be correlated with broad leaves and dilated, rounded and perfoliate leaf bases. Furthermore, two distinct subgroups were shown due to different number of vallecular vittae in each furrow, which may be applied as a basic classification factor for further investigations on the genus Bupleurum.
In this study, the fruit anatomy of all Bupleurum species in Iran, except B. wolffianum due to lack of available specimens, were studied for the first time. Some of the most taxonomically important fruit characteristics are as follows: mericarp shape in transverse section, characteristic of ribs, ratio of mericarp width to thickness, existence of oil ducts, quantity of vallecular and commissural vittae, ratio of height to width of dorsal rib, and shape of endocarp in transverse section. Important characters for each species are described and an identification key is provided to portray the similarities and relationships between the species. Anatomical studies also confirmed the anatomical and morphological differences between Sect. Perfoliata and other members of the genus. Absence of vittae in Sect. perfoliata may be correlated with broad leaves and dilated, rounded and perfoliate leaf bases. Furthermore, two distinct subgroups were shown due to different number of vallecular vittae in each furrow, which may be applied as a basic classification factor for further investigations on the genus Bupleurum.