<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
The periorbital morphology of goniopholidids is discussed, exploring the diversity of patterns an... more The periorbital morphology of goniopholidids is discussed, exploring the diversity of patterns and the relevance of the data for phylogenetic studies. Revision of material is focused on Goniopholis spp. and aff. Goniopholis spp., from England, Germany, and Belgium, providing a comparative description of their interorbital morphology. Traditional interpretation of the interorbital elements in species of Goniopholis (G. simus, G. baryglyphaeus),
ABSTRACT Metatetrapous valdensis Nopcsa, 1923 from the late Berriasian of northwestern Germany wa... more ABSTRACT Metatetrapous valdensis Nopcsa, 1923 from the late Berriasian of northwestern Germany was the first dinosaur ichnotaxon ever attributed to a thyreophoran trackmaker. However, the subsequent lost of the original material made this identification and the status of the ichnotaxon questionable for many subsequent authors. This situation was aggravated by the fact that there are only very brief original descriptions accompanied by a single drawing. A reconsideration of the original description recognizes M. valdensis as a valid ichnotaxon, which, albeit showing great resemblance in pes morphology to similar ichnotaxa, stands out from them by a tetradactyl manus. It not only holds its original systematic attribution, but also has sparked early hypotheses on the phylogeny of dinosaurs already in 1922, possibly for the first time based upon tracks. Two surviving natural hypichnial casts of ankylosaurian pes imprints from the same stratum cannot be straightforwardly identified with the type material due to a lack of documentation. However, comprehensive circumstantial evidence, including complete accordance in size and morphology among others, strongly supports such an association. The tracks confirm the presence of ankylosaurs in this lacustrine-deltaic setting as a very rare element of the local dinosaur fauna.
The Saxonian Cretaceous Basin constitutes an important source of rare Late Cretaceous marine amni... more The Saxonian Cretaceous Basin constitutes an important source of rare Late Cretaceous marine amniote fossils from Germany. It is also historically famous, having been documented in a series of monographic works published by the distinguished German palaeontologist Hanns Bruno Geinitz in the nineteenth century. The most productive rock units include the upper Cenomanian Dölzschen Formation and upper Turonian Strehlen and Weinböhla limestones (lower Strehlen Formation). A survey of curated specimens recovered from these deposits has now identified isolated teeth of probable polycotylid and elasmosaurid plesiosaurians, as well as several humeri that are referred to protostegid marine turtles. The Saxonian Cretaceous Basin formed a continuous epeiric seaway with the Bohemian Cretaceous Basin during late Cenomanian – Turonian time. A western connection to the North Sea Basin also existed via the North German and Münsterland Cretaceous basins. The Mesozoic marine amniote remains from thes...
Metriorhynchidae was a clade of extinct crocodylomorphs that adapted to a pelagic lifestyle, beco... more Metriorhynchidae was a clade of extinct crocodylomorphs that adapted to a pelagic lifestyle, becoming a key component of Mesozoic lagoonal and coastal marine ecosystems. The type genus Metriorhynchus is one of the best-known genera of Mesozoic crocodylomorphs, and since the mid-19th century, the ‘concept’ of Metriorhynchus has become associated with the referred species Me. superciliosus. Historically Metriorhynchus has been the most species-rich genus in Metriorhynchidae, with most Middle Jurassic species and many Late Jurassic species referred to the genus at some point in their history. However, the type species Me. geoffroyii has largely been omitted in the literature. Its type series is a chimera of multiple metriorhynchid species, and a type specimen has never been designated. Moreover, phylogenetic analyses have repeatedly shown that the 19th–20th century concept of Metriorhynchus is not monophyletic – to the point where only referring every metriorhynchid species, and some b...
Alcheringa: An Australasian Journal of Palaeontology, 2017
Sachs, S., Hornung, J.J., Lallensack, J.N. & Kear, B.P., November.2017. First evidence of a l... more Sachs, S., Hornung, J.J., Lallensack, J.N. & Kear, B.P., November.2017. First evidence of a large predatory plesiosaurian from the Lower Cretaceous non-marine ‘Wealden facies’ deposits of northwestern Germany. Alcheringa 42, 501-508. ISSN 0311-5518. Here, we describe the incomplete mandible of a large-skulled ‘pliosauromorph’ plesiosaurian from the Lower Cretaceous (Berriasian) of northwestern Germany. The fossil derives from limnic–brackish ‘Wealden facies’ deposits of the Deister Formation (Bückeberg Group), and is preserved as a natural mould in fine-grained sandstone. Examination of the original remains, in conjunction with a three-dimensional photogrammetrically digitized ‘cast’, revealed a conspicuous rosette of symphyseal alveoli, which would otherwise typically characterize Early–Middle Jurassic macrophagous plesiosaurians including rhomaleosaurids and the pliosaurid Simolestes. The Deister Formation ‘pliosauromorph’ represents the first record of a large-bodied plesiosaurian macrocarnivore from the ‘Wealden-facies’ strata of Europe, and thus adds a previously unrecognized trophic level of aquatic apex predators to the Early Cretaceous non-marine ecosystems of Europe. Sven Sachs* [sachs.pal@gmail.com] Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany and Im Hof 9, 51766 Engelskirchen, Germany; Jahn J. Hornung [jahn.hornung@yahoo.de] Niedersächsisches Landesmuseum Hannover, Willy-Brandt-Allee 5, 30169 Hannover, Germany; Jens N. Lallensack [jens.lallensack@uni-bonn.de] Steinmann-Institut für Geologie, Mineralogie und Paläontologie, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden.
In 1908, Hermann Löns outlined the concept of a Quintär” (“Quintary period”) to describe geologic... more In 1908, Hermann Löns outlined the concept of a Quintär” (“Quintary period”) to describe geological and biological manifestations of the Age of Humans. His definition of the “Quintary” consisted of twocomponents: a lithostratigraphical (“Quintary deposits” or “Quintary stratum”), and a faunistic (“Quintary fauna”) one. With a view on the stratigraphical component, Löns already anticipated the approach of the 21st Century to establish a geological definition for the Age of Humansbased on physical manifestations and geological evidence, that is nowadays reflected in the attempts to define the anthropocene. Transposed into modern terminology, the “Quintary stratum” is definedas a lithostratigraphical unit (Quintary Lithosome), that contains all deposits in which non-anthropogenic sedimentary processes have been replaced or modified by anthropogenic or technogenic activity. The Quintary Lithosome has a diachronous base and intercalates in its lower part vertically and laterally with non...
Based on a detailed morphological comparison of the original figures, the lost holotype of “Ornit... more Based on a detailed morphological comparison of the original figures, the lost holotype of “Ornithocheirus hilsensis” is identified as the distal part of the proximal pedal phalanx from digit I of a large-sized theropod. The distinctness in the morphology of the distal epiphysis of this element from that present in the manus and in pedal digits II-IV of most theropods may have contributed to the ambiguous interpretation of this specimen in the course of discussion since the 1880s. Features that have been interpreted as indicating pneumaticity – that would support a pterosaur affiliation – can be alternatively explained by taphonomic and diagenetic processes. Aside of this unresolved question, the published information do not indicate the presence of any pterosaur synapomorphies. Although clearly a nomen dubium, “Ornithocheirus hilsensis” is a precious record of a large-sized theropod near the Valanginian/Hauterivian boundary of Central Europe. It is furthermore of significance as on...
<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
The periorbital morphology of goniopholidids is discussed, exploring the diversity of patterns an... more The periorbital morphology of goniopholidids is discussed, exploring the diversity of patterns and the relevance of the data for phylogenetic studies. Revision of material is focused on Goniopholis spp. and aff. Goniopholis spp., from England, Germany, and Belgium, providing a comparative description of their interorbital morphology. Traditional interpretation of the interorbital elements in species of Goniopholis (G. simus, G. baryglyphaeus),
ABSTRACT Metatetrapous valdensis Nopcsa, 1923 from the late Berriasian of northwestern Germany wa... more ABSTRACT Metatetrapous valdensis Nopcsa, 1923 from the late Berriasian of northwestern Germany was the first dinosaur ichnotaxon ever attributed to a thyreophoran trackmaker. However, the subsequent lost of the original material made this identification and the status of the ichnotaxon questionable for many subsequent authors. This situation was aggravated by the fact that there are only very brief original descriptions accompanied by a single drawing. A reconsideration of the original description recognizes M. valdensis as a valid ichnotaxon, which, albeit showing great resemblance in pes morphology to similar ichnotaxa, stands out from them by a tetradactyl manus. It not only holds its original systematic attribution, but also has sparked early hypotheses on the phylogeny of dinosaurs already in 1922, possibly for the first time based upon tracks. Two surviving natural hypichnial casts of ankylosaurian pes imprints from the same stratum cannot be straightforwardly identified with the type material due to a lack of documentation. However, comprehensive circumstantial evidence, including complete accordance in size and morphology among others, strongly supports such an association. The tracks confirm the presence of ankylosaurs in this lacustrine-deltaic setting as a very rare element of the local dinosaur fauna.
The Saxonian Cretaceous Basin constitutes an important source of rare Late Cretaceous marine amni... more The Saxonian Cretaceous Basin constitutes an important source of rare Late Cretaceous marine amniote fossils from Germany. It is also historically famous, having been documented in a series of monographic works published by the distinguished German palaeontologist Hanns Bruno Geinitz in the nineteenth century. The most productive rock units include the upper Cenomanian Dölzschen Formation and upper Turonian Strehlen and Weinböhla limestones (lower Strehlen Formation). A survey of curated specimens recovered from these deposits has now identified isolated teeth of probable polycotylid and elasmosaurid plesiosaurians, as well as several humeri that are referred to protostegid marine turtles. The Saxonian Cretaceous Basin formed a continuous epeiric seaway with the Bohemian Cretaceous Basin during late Cenomanian – Turonian time. A western connection to the North Sea Basin also existed via the North German and Münsterland Cretaceous basins. The Mesozoic marine amniote remains from thes...
Metriorhynchidae was a clade of extinct crocodylomorphs that adapted to a pelagic lifestyle, beco... more Metriorhynchidae was a clade of extinct crocodylomorphs that adapted to a pelagic lifestyle, becoming a key component of Mesozoic lagoonal and coastal marine ecosystems. The type genus Metriorhynchus is one of the best-known genera of Mesozoic crocodylomorphs, and since the mid-19th century, the ‘concept’ of Metriorhynchus has become associated with the referred species Me. superciliosus. Historically Metriorhynchus has been the most species-rich genus in Metriorhynchidae, with most Middle Jurassic species and many Late Jurassic species referred to the genus at some point in their history. However, the type species Me. geoffroyii has largely been omitted in the literature. Its type series is a chimera of multiple metriorhynchid species, and a type specimen has never been designated. Moreover, phylogenetic analyses have repeatedly shown that the 19th–20th century concept of Metriorhynchus is not monophyletic – to the point where only referring every metriorhynchid species, and some b...
Alcheringa: An Australasian Journal of Palaeontology, 2017
Sachs, S., Hornung, J.J., Lallensack, J.N. & Kear, B.P., November.2017. First evidence of a l... more Sachs, S., Hornung, J.J., Lallensack, J.N. & Kear, B.P., November.2017. First evidence of a large predatory plesiosaurian from the Lower Cretaceous non-marine ‘Wealden facies’ deposits of northwestern Germany. Alcheringa 42, 501-508. ISSN 0311-5518. Here, we describe the incomplete mandible of a large-skulled ‘pliosauromorph’ plesiosaurian from the Lower Cretaceous (Berriasian) of northwestern Germany. The fossil derives from limnic–brackish ‘Wealden facies’ deposits of the Deister Formation (Bückeberg Group), and is preserved as a natural mould in fine-grained sandstone. Examination of the original remains, in conjunction with a three-dimensional photogrammetrically digitized ‘cast’, revealed a conspicuous rosette of symphyseal alveoli, which would otherwise typically characterize Early–Middle Jurassic macrophagous plesiosaurians including rhomaleosaurids and the pliosaurid Simolestes. The Deister Formation ‘pliosauromorph’ represents the first record of a large-bodied plesiosaurian macrocarnivore from the ‘Wealden-facies’ strata of Europe, and thus adds a previously unrecognized trophic level of aquatic apex predators to the Early Cretaceous non-marine ecosystems of Europe. Sven Sachs* [sachs.pal@gmail.com] Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany and Im Hof 9, 51766 Engelskirchen, Germany; Jahn J. Hornung [jahn.hornung@yahoo.de] Niedersächsisches Landesmuseum Hannover, Willy-Brandt-Allee 5, 30169 Hannover, Germany; Jens N. Lallensack [jens.lallensack@uni-bonn.de] Steinmann-Institut für Geologie, Mineralogie und Paläontologie, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussallee 8, 53115 Bonn, Germany; Benjamin P. Kear [benjamin.kear@em.uu.se] Museum of Evolution, Uppsala University, Norbyvägen 16, Uppsala SE-752 36, Sweden.
In 1908, Hermann Löns outlined the concept of a Quintär” (“Quintary period”) to describe geologic... more In 1908, Hermann Löns outlined the concept of a Quintär” (“Quintary period”) to describe geological and biological manifestations of the Age of Humans. His definition of the “Quintary” consisted of twocomponents: a lithostratigraphical (“Quintary deposits” or “Quintary stratum”), and a faunistic (“Quintary fauna”) one. With a view on the stratigraphical component, Löns already anticipated the approach of the 21st Century to establish a geological definition for the Age of Humansbased on physical manifestations and geological evidence, that is nowadays reflected in the attempts to define the anthropocene. Transposed into modern terminology, the “Quintary stratum” is definedas a lithostratigraphical unit (Quintary Lithosome), that contains all deposits in which non-anthropogenic sedimentary processes have been replaced or modified by anthropogenic or technogenic activity. The Quintary Lithosome has a diachronous base and intercalates in its lower part vertically and laterally with non...
Based on a detailed morphological comparison of the original figures, the lost holotype of “Ornit... more Based on a detailed morphological comparison of the original figures, the lost holotype of “Ornithocheirus hilsensis” is identified as the distal part of the proximal pedal phalanx from digit I of a large-sized theropod. The distinctness in the morphology of the distal epiphysis of this element from that present in the manus and in pedal digits II-IV of most theropods may have contributed to the ambiguous interpretation of this specimen in the course of discussion since the 1880s. Features that have been interpreted as indicating pneumaticity – that would support a pterosaur affiliation – can be alternatively explained by taphonomic and diagenetic processes. Aside of this unresolved question, the published information do not indicate the presence of any pterosaur synapomorphies. Although clearly a nomen dubium, “Ornithocheirus hilsensis” is a precious record of a large-sized theropod near the Valanginian/Hauterivian boundary of Central Europe. It is furthermore of significance as on...
<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
<p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontoge... more <p>Supplementary data (PAST-output) to Hornung et al.:</p> <p>Diversity, ontogeny, or both? A morphometric approach to iguanodontian ornithopod (Dinosauria: Ornithischia) track assemblages from the Berriasian (Lower Cretaceous) of northwestern Germany</p> <p>- 2D Semi-landmarks dataset</p> <p>- Previous dataset with procrustes fitting</p
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