^Branstetter, Michael G.; Danforth, Bryan N.; Pitts, James P.; Faircloth, Brant C.; Ward, Philip S.; Buffington, Matthew L.; Gates, Michael W.; Kula, Robert R. et al. (2017). “Phylogenomic Insights into the Evolution of Stinging Wasps and the Origins of Ants and Bees”. Current Biology 27 (7): 1019–1025. doi:10.1016/j.cub.2017.03.027. PMID 28376325
^S.A. Marshall 2023 Hymenoptera: The Natural History and Diversity of Wasps, Bees and Ants. Firefly Books
