Christian Ammer

ABSTRACT Context Identifying the traits of exotic species may explain their invasiveness and help control the negative impacts of these species on native ecosystems. In this study, we investigated root competition as an important driver for the competitiveness of two exotic tree species in the seedling stage. Aims In a pot experiment, carried out in Central Europe, we studied the effect of root competition of two exotic tree species (Prunus serotina Ehrh. and Robinia pseudoacacia L.) on biomass allocation of two native ones (Quercus robur L. and Carpinus betulus L.). Methods Seedlings of these species were exposed to intra- and interspecific mixtures with and without the effect of root competition. For this, belowground plastic partitions were installed in the pots. Results Competition intensity in the pots increased over time, irrespective of mixture type. However, this increase was much higher for the native species if mixed with the exotic species compared to monocultures or inter-native mixtures. In addition to ontogeny, competition affected biomass allocation patterns. Under root competition by the exotic species, the native species preferentially allocated their biomass to the roots. A higher allocation to the roots was mainly achieved at the expense of leaf and branch biomass. Conclusion Root competition of P. serotina and R. pseudoacacia may be a reason for the lack of Q. robur and C. betulus in the seedling stage in natural environments where all four species occur.

ABSTRACT Predicted intense and prolonged drought events challenge forest management. Thinning is debated as a silvicultural measure for reducing drought risk in densely established forest stands. We report on a thinning experiment in a 26-year-old Norway spruce stand (Picea abies), comprising of two thinning intensities and one unthinnned control. The removal of 43% (moderate thinning, MT) and 67% (heavy thinning, HT) of the initial basal area led to increased water availability during the entire three year observation period. Stand-level transpiration (Es) was decreased by about 25% upon moderate, and by about 50% upon heavy thinning during the first year after the interventions had been carried out. However, differences in Es across the treatments decreased within three years after thinning mainly due to increased single-tree transpiration and additional understory evapotranspiration at HT. Nevertheless, due to lower interception and transpiration on the thinned plots three years after treatment MT and HT still showed a substantial surplus in extractable soil water. The results showed that the main determinants concerning the extent of the mitigation effect with increasing thinning intensity were the available soil water storage capacity and the emerging understory vegetation. We conclude that repeated moderate thinning, through enhancing the water availability to the remaining trees, can mitigate drought risk in young spruce stands and thus, represent a viable silvicultural measure in anticipating possible water limitations due to climate change.

We used height growth data from a 7-year field experiment with European beech (Fagus sylvatica L.) seedlings to test the hypothesis that the effects of above- and belowground resources on height growth depend on seedling size and age. Photosynthetically active radiation (PAR) was determined by hemispheric photography, and estimates of fine root biomass of the overstory trees were used as an inverse proportional surrogate for belowground resource availability. For recently germinated seedlings growing under the canopy of Picea abies (L.) Karst., belowground resource availability affected height growth more than light. During subsequent stages of seedling development, apart from initial seedling size, PAR increasingly determined seedling growth. Besides initial size, seedling age determined the effects of above- and belowground resources on seedling height growth. In seedlings identical in initial size but differing in age, the increase in height growth with increasing PAR was greater...

ABSTRACT With an increasing demand of sustainable raw materials for bioenergy use, coppicing as management approach to increase the biomass production of forests is becoming of greater importance. This study describes the parameterization of biomass equations for six tree species traditionally used in coppices forests, namely sycamore maple (Acer pseudoplatanus L.), field maple (Acer campestre L.), European ash (Fraxinus excelsior L.), European hornbeam (Carpinus betulus L.), downy birch (Betula pubescens Ehrh.), and common hazel (Corylus avellana L.) growing in coppice-with-standard systems in Lower Saxony, Germany. The parameterization was based on measurements of over 950 trees sampled from two forest sites. The sampled trees were felled and separated into three biomass compartments (stem, coarse branches, and fine brushwood) and weighed on site. The dry weight of sub samples from each compartment was measured. Equations were derived for total aboveground biomass, stem biomass, and crown biomass using regression analyses. We either used diameter at breast height as single independent explanatory variable or in combination with tree height. Biomass production of stump sprouts and generatively grown stems was compared for ash and sycamore maple. In the same age classes, it was found that ash stump sprouts had a slightly higher production than seed-grown stems. For sycamore maple, no difference was detected.

ABSTRACT We assessed the effectiveness of different management strategies against the non-native invasive tree species black cherry (Prunus serotina Ehrh.). The species causes substantial management problems in European forest ecosystems, like the Valle del Ticino Biosphere Reserve in Italy, by suppressing the regeneration of native tree species. This can modify ecological key processes and cause biodiversity loss. Since chemical and biological control has mainly been abandoned in European forest ecosystem management, mechanical control measures are presently the preferred option to proceed against the black cherry but have shown very limited results in the reserve. The aim was to control the success of felling the species and to test other mechanical control methods such as girdling and snapping the trees with regard to their efficiency by quantifying the species' growth reactions. For this purpose, observational studies were conducted in two forest stands of which one was treated in 1996 and the other more recently in 2009. A subsample of resprouting stumps was treated a second time in 2010 to observe the effect of a direct second cutback. An experimental study was implemented in a third forest stand also in 2010 to compare three different mechanical control methods. The results suggest that felling black cherry is ineffective if the objective is to reduce the species' abundance because resprouts occur on 100% of the treated trees and biomass increment is not reduced in the long term. Girdling proved to be the most effective treatment across the diameter classes considered.

Abstract We compared different approaches for model-ling height growth of individual beech seedlings in a controlled factorial experiment as well as in field data from naturally regenerated beech seedlings under the canopy of overstorey mature beech trees. Several competition ...

For several reasons the conversion of mono-species into mixed-species forests is presently a major concern of forest management and policy in Central Europe. Although it is possible to show a clear trend in favour of mixed-species forests, private forest owners and some forest economists have often not favoured mixed forests, assuming that they are less profitable. The trend towards mixed