The dispersal of Homo sapiens across Eurasia during MIS 3 in the Late Pleistocene is marked by te... more The dispersal of Homo sapiens across Eurasia during MIS 3 in the Late Pleistocene is marked by technological shifts and other behavioral changes, known in the archaeological record under the term of Initial Upper Paleolithic (IUP). Bacho Kiro Cave in north Bulgaria, re-excavated by us from 2015 to 2021, is one of the reference sites for this phenomenon. The newly excavated lithic assemblages dated by radiocarbon between 45,040 and 43,280 cal BP and attributed to Homo sapiens encompass more than two thousand lithic artifacts. The lithics, primarily from Layer N1-I, exist amid diverse fauna remains, human fossils, pierced animal teeth pendants, and sediment with high organic content. This article focuses on the technological aspects of the IUP lithics, covering raw material origin and use-life, blank production, on-site knapping activities, re-flaking of lithic implements, and the state of retouched lithic components. We apply petrography for the identification of silicites and other used stones. We employ chaîne opératoire and reduction sequence approaches to profile the lithics techno-typologically and explore the lithic economy, particularly blade production methods, knapping techniques, and artifact curation. Raw material analysis reveals Lower Cretaceous flints from Ludogorie and Upper Cretaceous flints from the Danube region, up to 190 km and 130 km, respectively, from Bacho Kiro Cave, indicating long-distance mobility and finished products transport. Imported lithic implements, were a result of unidirectional and bidirectional non-Levallois laminar technology, likely of volumetric concept. Systematic on-anvil techniques (bipolar knapping) and tool segmentation indicate re-flaking and reshaping of lithic implements, reflecting on-site curation and multifaceted lithic economy. A limited comparison with other IUP sites reveals certain shared features and also regional variations. Bacho Kiro Cave significantly contributes to understanding the technological and behavioral evolution of early Homo sapiens in western Eurasia.
Artefacts made from stones, bones and teeth are fundamental to our understanding of human subsist... more Artefacts made from stones, bones and teeth are fundamental to our understanding of human subsistence strategies, behaviour and culture in the Pleistocene. Although these resources are plentiful, it is impossible to associate artefacts to specific human individuals1 who can be morphologically or genetically characterized, unless they are found within burials, which are rare in this time period. Thus, our ability to discern the societal roles of Pleistocene individuals based on their biological sex or genetic ancestry is limited2–5. Here we report the development of a non-destructive method for the gradual release of DNA trapped in ancient bone and tooth artefacts. Application of the method to an Upper Palaeolithic deer tooth pendant from Denisova Cave, Russia, resulted in the recovery of ancient human and deer mitochondrial genomes, which allowed us to estimate the age of the pendant at approximately 19,000–25,000 years. Nuclear DNA analysis identifies the presumed maker or wearer o...
Bone surface modifications are crucial for understanding human subsistence and dietary behaviour,... more Bone surface modifications are crucial for understanding human subsistence and dietary behaviour, and can inform about the techniques employed in the production and use of bone tools. Permission to destructively sample such unique artefacts is not always granted. The recent development of non-destructive proteomic extraction techniques has provided some alternatives for the analysis of rare and culturally significant artefacts, including bone tools and personal ornaments. The Eraser Extraction Method (EEM), first developed for ZooMS analysis of parchment, has recently been applied to bone and ivory specimens. To test the potential impact of the EEM on ancient bone surfaces, we analyse six anthropogenically modified Palaeolithic bone specimens from Bacho Kiro Cave (Bulgaria) through a controlled sampling experiment using qualitative and 3D quantitative microscopy. Although the overall bone topography is generally preserved, our findings demonstrate a slight flattening of the microtop...
Abstract This paper reports the preliminary results of two test pit excavations in Toplitsa cave ... more Abstract This paper reports the preliminary results of two test pit excavations in Toplitsa cave in Northern Bulgaria and discusses the potential for future investigations of the Pleistocene layers associated with Marine Isotope Stage 3 and the transitional Middle to Upper Palaeolithic period. We describe the lithostratigraphic layers, archaeological finds and features attributable to the Upper Palaeolithic. The preliminary chrono-cultural interpretation of the early Upper Palaeolithic geoarchaeological deposit at Toplitsa cave is based on the identification of Campanian Ignimbrite/Y-5 (CI) tephra, radiocarbon dates and the technological assessment of a small lithic artefact assemblage in comparison with the regional Initial Upper Palaeolithic (IUP) technologies. Toplitsa cave, located in the Eastern Balkans (Northern Bulgaria), was excavated in 2011 with the main goal of investigating deposits encompassing the transition from the Middle to Upper Palaeolithic. Two test pits (TP) were excavated: TP I, incorporating four lithostratigraphic layers, was located in the cave interior and TP II, encompassing six lithostratigraphic layers, was established closer to the cave entrance. Both TPs were generally poor in archaeological remains which impacted the chrono-cultural attribution of the Pleistocene layers. Moreover, in some spots Holocene faunal remains likely intruded into older layers. In consequence, the preliminary chronological investigation has been focused on applying both tephrochronology and radiocarbon dating of the early Upper Palaeolithic deposits. We present a preliminary assessment of the stratigraphy and report on the identification of large amounts of glass shards belonging to the CI eruption (ca. 39,000 years old (ka BP)) in TP II Layers 2 and 3, below which there is a well preserved hearth. This fire-pit carved in Layer 5 (TP II) and underlying the Y-5 tephra is the most significant find. Two Ursus spelaeus teeth from the underlying Layer 5 have been radiocarbon dated to 49,770–45,750 calibrated years before present (cal BP; 95.4% probability) and to 48,210–45,050 cal BP (95.4% probability). The lithics found in Layer 5 are technologically consistent with the Initial Upper Palaeolithic (IUP) or possibly with other transitional Middle to Upper Palaeolithic assemblages. Layer 5 from TP II in Toplitsa overlaps chronologically with other regional records. These include the final Middle Palaeolithic with leaf points from Musselievo (estimated to ca. 50–45 ka cal BP), the transitional Middle to Upper Palaeolithic assemblage from Temnata Cave Layer VI (trench II, which is covered by CI tephra and has been dated to older than 39 ka BP), the earliest laminar Upper Palaeolithic (i.e. IUP) from Temnata Cave Layer 4 (trench I, dating from 47.6 to 40.5 ka cal BP), the IUP from Bacho Kiro Cave Layers J and I (dating to 46.0–43.3 ka cal BP) and Kozarnika cave Layer 6/7 (dated to ca. 49.1–44.0 ka cal BP).
The behavioral dynamics underlying the expansion of Homo sapiens into Europe remains a crucial to... more The behavioral dynamics underlying the expansion of Homo sapiens into Europe remains a crucial topic in human evolution. Owing to poor bone preservation, past studies have strongly focused on the Initial Upper Paleolithic (IUP) stone tool record. Recent excavations and extensive radiocarbon dating at Bacho Kiro Cave (Bulgaria) pushed back the arrival of IUP H. sapiens into Europe to ca. 45,000 years ago. This site has exceptional bone preservation, and we present the study of 7431 faunal remains from across two IUP layers (I and J) and one Middle Paleolithic layer (K). We identified a shift in site use and occupation intensity through time, marked by increased find density and human modifications in Layer I. Alongside a decrease in carnivore presence and seasonality data demonstrating human presence in all seasons, this indicates a more frequent or prolonged occupation of the site by IUP groups. Contrarily, the dietary focus across the IUP and Middle Paleolithic layers is similar, centered on the exploitation of species from a range of habitats including Bos/Bison, Cervidae, Equidae, and Caprinae. While body parts of large herbivores were selectively transported into the site, the bear remains suggest that these animals died in the cave itself. A distinct aspect of the IUP occupation is an increase in carnivore remains with human modifications, including these cave bears but also smaller taxa (e.g., Canis lupus, Vulpes vulpes). This can be correlated with their exploitation for pendants, and potentially for skins and furs. At a broader scale, we identified similarities in subsistence behavior across IUP sites in Europe and western Asia. It appears that the first IUP occupations were less intense with find densities and human modifications increasing in succeeding IUP layers. Moreover, the exploitation of small game appears to be limited across IUP sites, while carnivore exploitation seems a recurrent strategy.
The dispersal of Homo sapiens across Eurasia during MIS 3 in the Late Pleistocene is marked by te... more The dispersal of Homo sapiens across Eurasia during MIS 3 in the Late Pleistocene is marked by technological shifts and other behavioral changes, known in the archaeological record under the term of Initial Upper Paleolithic (IUP). Bacho Kiro Cave in north Bulgaria, re-excavated by us from 2015 to 2021, is one of the reference sites for this phenomenon. The newly excavated lithic assemblages dated by radiocarbon between 45,040 and 43,280 cal BP and attributed to Homo sapiens encompass more than two thousand lithic artifacts. The lithics, primarily from Layer N1-I, exist amid diverse fauna remains, human fossils, pierced animal teeth pendants, and sediment with high organic content. This article focuses on the technological aspects of the IUP lithics, covering raw material origin and use-life, blank production, on-site knapping activities, re-flaking of lithic implements, and the state of retouched lithic components. We apply petrography for the identification of silicites and other used stones. We employ chaîne opératoire and reduction sequence approaches to profile the lithics techno-typologically and explore the lithic economy, particularly blade production methods, knapping techniques, and artifact curation. Raw material analysis reveals Lower Cretaceous flints from Ludogorie and Upper Cretaceous flints from the Danube region, up to 190 km and 130 km, respectively, from Bacho Kiro Cave, indicating long-distance mobility and finished products transport. Imported lithic implements, were a result of unidirectional and bidirectional non-Levallois laminar technology, likely of volumetric concept. Systematic on-anvil techniques (bipolar knapping) and tool segmentation indicate re-flaking and reshaping of lithic implements, reflecting on-site curation and multifaceted lithic economy. A limited comparison with other IUP sites reveals certain shared features and also regional variations. Bacho Kiro Cave significantly contributes to understanding the technological and behavioral evolution of early Homo sapiens in western Eurasia.
Artefacts made from stones, bones and teeth are fundamental to our understanding of human subsist... more Artefacts made from stones, bones and teeth are fundamental to our understanding of human subsistence strategies, behaviour and culture in the Pleistocene. Although these resources are plentiful, it is impossible to associate artefacts to specific human individuals1 who can be morphologically or genetically characterized, unless they are found within burials, which are rare in this time period. Thus, our ability to discern the societal roles of Pleistocene individuals based on their biological sex or genetic ancestry is limited2–5. Here we report the development of a non-destructive method for the gradual release of DNA trapped in ancient bone and tooth artefacts. Application of the method to an Upper Palaeolithic deer tooth pendant from Denisova Cave, Russia, resulted in the recovery of ancient human and deer mitochondrial genomes, which allowed us to estimate the age of the pendant at approximately 19,000–25,000 years. Nuclear DNA analysis identifies the presumed maker or wearer o...
Bone surface modifications are crucial for understanding human subsistence and dietary behaviour,... more Bone surface modifications are crucial for understanding human subsistence and dietary behaviour, and can inform about the techniques employed in the production and use of bone tools. Permission to destructively sample such unique artefacts is not always granted. The recent development of non-destructive proteomic extraction techniques has provided some alternatives for the analysis of rare and culturally significant artefacts, including bone tools and personal ornaments. The Eraser Extraction Method (EEM), first developed for ZooMS analysis of parchment, has recently been applied to bone and ivory specimens. To test the potential impact of the EEM on ancient bone surfaces, we analyse six anthropogenically modified Palaeolithic bone specimens from Bacho Kiro Cave (Bulgaria) through a controlled sampling experiment using qualitative and 3D quantitative microscopy. Although the overall bone topography is generally preserved, our findings demonstrate a slight flattening of the microtop...
Abstract This paper reports the preliminary results of two test pit excavations in Toplitsa cave ... more Abstract This paper reports the preliminary results of two test pit excavations in Toplitsa cave in Northern Bulgaria and discusses the potential for future investigations of the Pleistocene layers associated with Marine Isotope Stage 3 and the transitional Middle to Upper Palaeolithic period. We describe the lithostratigraphic layers, archaeological finds and features attributable to the Upper Palaeolithic. The preliminary chrono-cultural interpretation of the early Upper Palaeolithic geoarchaeological deposit at Toplitsa cave is based on the identification of Campanian Ignimbrite/Y-5 (CI) tephra, radiocarbon dates and the technological assessment of a small lithic artefact assemblage in comparison with the regional Initial Upper Palaeolithic (IUP) technologies. Toplitsa cave, located in the Eastern Balkans (Northern Bulgaria), was excavated in 2011 with the main goal of investigating deposits encompassing the transition from the Middle to Upper Palaeolithic. Two test pits (TP) were excavated: TP I, incorporating four lithostratigraphic layers, was located in the cave interior and TP II, encompassing six lithostratigraphic layers, was established closer to the cave entrance. Both TPs were generally poor in archaeological remains which impacted the chrono-cultural attribution of the Pleistocene layers. Moreover, in some spots Holocene faunal remains likely intruded into older layers. In consequence, the preliminary chronological investigation has been focused on applying both tephrochronology and radiocarbon dating of the early Upper Palaeolithic deposits. We present a preliminary assessment of the stratigraphy and report on the identification of large amounts of glass shards belonging to the CI eruption (ca. 39,000 years old (ka BP)) in TP II Layers 2 and 3, below which there is a well preserved hearth. This fire-pit carved in Layer 5 (TP II) and underlying the Y-5 tephra is the most significant find. Two Ursus spelaeus teeth from the underlying Layer 5 have been radiocarbon dated to 49,770–45,750 calibrated years before present (cal BP; 95.4% probability) and to 48,210–45,050 cal BP (95.4% probability). The lithics found in Layer 5 are technologically consistent with the Initial Upper Palaeolithic (IUP) or possibly with other transitional Middle to Upper Palaeolithic assemblages. Layer 5 from TP II in Toplitsa overlaps chronologically with other regional records. These include the final Middle Palaeolithic with leaf points from Musselievo (estimated to ca. 50–45 ka cal BP), the transitional Middle to Upper Palaeolithic assemblage from Temnata Cave Layer VI (trench II, which is covered by CI tephra and has been dated to older than 39 ka BP), the earliest laminar Upper Palaeolithic (i.e. IUP) from Temnata Cave Layer 4 (trench I, dating from 47.6 to 40.5 ka cal BP), the IUP from Bacho Kiro Cave Layers J and I (dating to 46.0–43.3 ka cal BP) and Kozarnika cave Layer 6/7 (dated to ca. 49.1–44.0 ka cal BP).
The behavioral dynamics underlying the expansion of Homo sapiens into Europe remains a crucial to... more The behavioral dynamics underlying the expansion of Homo sapiens into Europe remains a crucial topic in human evolution. Owing to poor bone preservation, past studies have strongly focused on the Initial Upper Paleolithic (IUP) stone tool record. Recent excavations and extensive radiocarbon dating at Bacho Kiro Cave (Bulgaria) pushed back the arrival of IUP H. sapiens into Europe to ca. 45,000 years ago. This site has exceptional bone preservation, and we present the study of 7431 faunal remains from across two IUP layers (I and J) and one Middle Paleolithic layer (K). We identified a shift in site use and occupation intensity through time, marked by increased find density and human modifications in Layer I. Alongside a decrease in carnivore presence and seasonality data demonstrating human presence in all seasons, this indicates a more frequent or prolonged occupation of the site by IUP groups. Contrarily, the dietary focus across the IUP and Middle Paleolithic layers is similar, centered on the exploitation of species from a range of habitats including Bos/Bison, Cervidae, Equidae, and Caprinae. While body parts of large herbivores were selectively transported into the site, the bear remains suggest that these animals died in the cave itself. A distinct aspect of the IUP occupation is an increase in carnivore remains with human modifications, including these cave bears but also smaller taxa (e.g., Canis lupus, Vulpes vulpes). This can be correlated with their exploitation for pendants, and potentially for skins and furs. At a broader scale, we identified similarities in subsistence behavior across IUP sites in Europe and western Asia. It appears that the first IUP occupations were less intense with find densities and human modifications increasing in succeeding IUP layers. Moreover, the exploitation of small game appears to be limited across IUP sites, while carnivore exploitation seems a recurrent strategy.
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