Modul FisTan 1
Modul FisTan 1
Modul FisTan 1
1. INTRODUCTION
2. ENERGY
e- Compe-
tency
http:\\smtom.lecture.ub.ac.id
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LECTURE LAYOUT
1. INTRODUCTION 2. ENERGY
1. Definition 1. Definition
2. History Of Plant 2. Energy Of Organisms
Physiology 3. Law of Conservation of
3. How Plants Work Energy
4. Syllabus 4. State and Form of Energy
References 5. Electron Energy
6. Free Energy
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1.1 Definition
What is Plant Physiology?
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INTRODUCTION
What is Plant Physiology about ?
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2. Jan van Helmont
began the research of the process in the mid-
1600s when he carefully measured the mass
of the soil used by a plant and the mass of the
plant as it grew.
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3. John Woodward
(1 May 1665 – 25 April 1728), an
English naturalist, antiquarian and
geologist, published experiments
in 1699 on growth of spearmint
(Mentha spicata) in different
sources of water.
He found that plants grew much
better in water with soil added
than in distilled water.
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4. Stephen Hales, FRS (17 September
1677 – 4 January 1761), an English
physiologist, chemist and inventor
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5. Joseph PriestleyO2
a chemist and minister, discovered that when he isolated a
volume of air under an inverted jar, and burned a candle in it,
the candle would burn out very quickly, much before it ran out
of wax. He further discovered that a mouse could similarly
"injure" air. He then showed that the air that had been "injured"
by the candle and the mouse could be restored by a plant.
6. Jan Ingenhousz O2 produce d by plants +light
a court physician to the Austrian Empress, repeated Priestley's
experiments in 1778, . He discovered that it was the influence of
sun and light on the plant that could cause it to rescue a mouse
in a matter of hours.
7. Jean Senebier CO2 taken up by plants
a French pastor, showed in 1796, that CO2 was the "fixed" or
"injured" air and that it was taken up by plants in
photosynthesis.
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8. Nicolas-Théodore de Saussure CO2 +H2O
soon afterwards, showed that the increase in mass of the
plant as it grows could not be due only to uptake of CO2,
but also to the incorporation of water. Thus the basic
reaction by which photosynthesis is used to produce food
(such as glucose) was outlined.
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1.3 How Plants Work
Plant Level 6. Christae 18. Golgi body
Shoot 7. Nucleus 19. Peroxisomes
Leaves 8. Nuclear membrane 20. Microfilaments
Stem 9. Nucleolus 21. Microtubules
Reproductive Organ
10. Centrosome 22. Plasmodesmata
Root
11. Ribosome 23. Rough
Cellular Level 12. Chloroplast endoplasmic
1. Cell wall 13. Chlorophyll reticulum
2. Cell membrane 14. Stroma 24. Smooth
3. Cytoplasm endoplasmic
15. Thylakoid disk reticulum
4. Vacuole 16. Granum
5. Mitochondrion 17. Amyloplast
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Plant Level
• The Shoot System
• Above ground (usually)
• Elevates the plant above the
soil
• Many functions including:
• photosynthesis
• reproduction & dispersal
• food and water conduction
• The Root System
• Underground (usually)
• Anchor the plant in the soil
• Absorb water and nutrients
• Conduct water and nutrients
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Cellular Level
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1. Cell wall - a thick, rigid membrane that surrounds a plant cell. This
layer of cellulose fiber gives the cell most of its support and structure.
The cell wall also bonds with other cell walls to form the structure of
the plant.
2. Cell membrane - the thin layer of protein and fat that surrounds the
cell, but is inside the cell wall. The cell membrane is semipermeable,
allowing some substances to pass into the cell and blocking others.
3. Cytoplasm - the jellylike material outside the cell nucleus in which the
organelles are located.
4. Vacuole - a large, membrane-bound space within a plant cell that is
filled with fluid. Most plant cells have a single vacuole that takes up
much of the cell. It helps maintain the shape of the cell.
5. Mitochondrion - spherical to rod-shaped organelles with a double
membrane. The inner membrane is infolded many times, forming a
series of projections (called cristae). The mitochondrion converts the
energy stored in glucose into ATP (adenosine triphosphate) for the
cell.
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6. Christae - (singular crista) the multiply-folded inner membrane of a
cell's mitochondrion that are finger-like projections. The walls of the
cristae are the site of the cell's energy production (it is where ATP is
generated).
7. Nucleus - spherical body containing many organelles, including the
nucleolus. The nucleus controls many of the functions of the cell (by
controlling protein synthesis) and contains DNA (in chromosomes).
The nucleus is surrounded by the nuclear membrane
8. Nuclear membrane - the membrane that surrounds the nucleus.
9. Nucleolus - an organelle within the nucleus - it is where ribosomal RNA
is produced.
10. Centrosome - (also called the "microtubule organizing center") a small
body located near the nucleus - it has a dense center and radiating
tubules. The centrosomes is where microtubules are made. During cell
division (mitosis), the centrosome divides and the two parts move to
opposite sides of the dividing cell. Unlike the centrosomes in animal
cells, plant cell centrosomes do not have centrioles.
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11. Ribosome - small organelles composed of RNA-rich cytoplasmic
granules , approximately 60 percent RNA and 40 percent protein , that
are sites of protein synthesis. In eukaryotes, ribosomes are made of
four strands of RNA. In prokaryotes, they consist of three strands of
RNA.
12. Chloroplast - an elongated or disc-shaped organelle containing
chlorophyll. Photosynthesis (in which energy from sunlight is
converted into chemical energy - food) takes place in the chloroplasts.
13. Chlorophyll - chlorophyll is a molecule that can use light energy from
sunlight to turn water and carbon dioxide gas into sugar and oxygen
(this process is called photosynthesis). Chlorophyll is magnesium
based and is usually green.
14. Stroma - part of the chloroplasts in plant cells, located within the inner
membrane of chloroplasts, between the grana.
15. Thylakoid disk - thylakoid disks are disk-shaped membrane structures
in chloroplasts that contain chlorophyll. Chloroplasts are made up of
stacks of thylakoid disks; a stack of thylakoid disks is called a granum.
Photosynthesis (the production of ATP molecules from sunlight) takes
place on thylakoid disks.
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16. Granum - (plural grana) A stack of thylakoid disks within the chloroplast is
called a granum.
17. Amyloplast - an organelle in some plant cells that stores starch.
Amyloplasts are found in starchy plants like tubers and fruits.
18. Golgi body - (also called the golgi apparatus or golgi complex) a flattened,
layered, sac-like organelle that looks like a stack of pancakes and is located
near the nucleus. The golgi body packages proteins and carbohydrates into
membrane-bound vesicles for "export" from the cell.
19. Peroxisomes - Microbodies are a diverse group of organelles that are found
in the cytoplasm, roughly spherical and bound by a single membrane.
There are several types of microbodies but peroxisomes are the most
common.
20. Microfilaments - Microfilaments are solid rods made of globular proteins
called actin. These filaments are primarily structural in function and are an
important component of the cytoskeleton.
21. Microtubules - These straight, hollow cylinders are found throughout the
cytoplasm of all eukaryotic cells (prokaryotes don't have them) and carry
out a variety of functions, ranging from transport to structural support.
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22. Plasmodesmata - Plasmodesmata are small tubes that connect plant
cells to each other, providing living bridges between cells.
23. Rough endoplasmic reticulum - (rough ER) a vast system of
interconnected, membranous, infolded and convoluted sacks that are
located in the cell's cytoplasm (the ER is continuous with the outer
nuclear membrane). Rough ER is covered with ribosomes that give it a
rough appearance. Rough ER transport materials through the cell and
produces proteins in sacks called cisternae (which are sent to the Golgi
body, or inserted into the cell membrane).
24. Smooth endoplasmic reticulum - (smooth ER) a vast system of
interconnected, membranous, infolded and convoluted tubes that are
located in the cell's cytoplasm (the ER is continuous with the outer
nuclear membrane). The space within the ER is called the ER lumen.
Smooth ER transport materials through the cell. It contains enzymes
and produces and digests lipids (fats) and membrane proteins; smooth
ER buds off from rough ER, moving the newly-made proteins and lipids
to the Golgi body and membranes
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1.4 SYLLABUS
LECTURE TOPICS CONTENTS
Introduction
1. ENERGY
Energy
Light Reaction: Synthesis of NADPH
2. PHOTOSYNTESIS I
Light Reaction: Synthesis of ATP
Dark Reaction: C3 Plants
3. PHOTOSYNTESIS II
Dark Reaction: C4 & CAM Plants
4. PHOTOSYNTESIS III Factors affecting photosynthesis
Glycolysis, TCA Cycle & Terminal
5. RESPIRATION
Oxidation
6. TRANSPORT SYSTEM Xylem Transport, Transpiration
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References
1. Taiz, L. and Zeiger, E., 1991. Plant Physiology. The
Benjamin/Cummings Publishing Co., Inc., Redwood City,
California
2. Salisbury, F.B. and C.W. Ross. 1992. Plant Physiology
3. Salisbury, F.B. and Ross, C., 1969. Plant Physiology. Wadsworth
Publishing Co. Inc., Belmont, California
4. Bidwell,R.G.S. 1979. Plant Physiology. Mac. Millan. Publishing,
New York
5. Devlin, R.M. and F.H. Witham. 1983. Plant Physiology. The Towa
State University Press
6. Gardner,F.P.,R.B. Pearce and R.L. Mitchell.1985, Physiology of
crop plants
7. Hall.D.O. and K.K. Rao 1981. Photosynthesis, London
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ENERGI & PROSES FISIOLOGI TUMBUHAN
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ENERGI
Berdasarkan sumber energi yang digunakan
organisme hidup dibagi menjadi 4 kelompok:
1. Chemotrophs (senyawa organik)
2. Phototrophs (cahaya)
3. Chemoheterotrops (bahan organik +
karbon) Manusia dan Hewan
4. Photoautotrophs (radiasi matahari + CO2
(sebagai sumber karbon) Tanaman
Tingkat tinggi
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Dalam studi energi Hukum
Thermodinamika I yang menyatakan:
Energi dapat ditransfer dan ditransformasi,
tetapi tidak dapat diciptakan dan tidak
dapat dimusnahkan Dikenal : Hukum
Kekekalan Energi dan Massa
Energi Surya yang diterima tanaman
berubah bentuk dari energi
elektromagnetik menjadi zat kimia (energi
kimia) yaitu senyawa yang mengandung
energi tinggi dan reduktan
Energi Kimia ini digunakan untuk
mensintesis Karbohidrat yang merupakan
bahan dasar pembentukan senyawa2
penyusun sel, jaringan, organ dan
keseluruhan tubuh tanaman
Energi Surya pada akhirnya berubah
menjadi Energi Biomassa dalam
pertumbuhan tanaman
Jadi ORGANISME adalah pentransformasi
energi
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ATP
ADENOSINE TRIPHOSPHAT (ATP)
merupakan asam nukleat yang menyediakan
energi secara teratur dalam semua sel
ATP tersusun dari Adenin Ribose dan tiga
Phosphat
Suatu organisme yang sedang bekerja,
menggunakan ATP secara terus menerus
ATP adalah sumberdaya yang dapat
diperbaharui, yang dapat diregenerasi dengan
cara penambahan phosphat ke ADP
Siklus ATP bergerak dengan laju yang sangat
cepat
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Persamaan Einstein: E = m.c 2
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Konsep Umum Energi
Suatu benda karena kedudukannya mempunyai
energi tambahan di luar dari massa benda itu
sendiri
Batu yang jatuh dari ketinggian tertentu di atas
permukaan tanah ke tempat yang lebih rendah
massanya tidak berubah
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Energi Kinetik
Energi yang terlibat dalam pergerakan suatu benda
disebut: Energi Kinetik / Energi pergerakan Obyek
bergerak melakukan kerja dengan cara menggerakkan
benda lain
Orang menulis atau berlari
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Teori Kinetik
Berasal dari kata Kinetikos = membuat gerakan /
aktivitas
Merupakan landasan dasar dari pengetahuan
energi
Konsep ini menyatakan: Semua partikel (atom &
molekul) berada dalam gerakan konstan pada
semua tingkat temperatur di atas nol absolut
Fungsi daun sebagai tempat masuknya CO2
melalui proses difusi
Energi radiasi matahari berubah menjadi energi
kimia dalam tanaman
Pergerakan suatu benda / obyek berdasarkan Teori
Kinetik dibedakan menjadi 3:
1. Gerak lurus (Kinetik Translasi)
2. Gerak berputar (Kinetik Rotasi)
3. Gerak bergetar (Kinetik Vibrasi)
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Catatan:
1. Suatu gas tersusun dari sejumlah besar
atom & molekul yang dipisahkan oleh jarak
yang relatif cukup besar dibandingkan
dengan ukurannya
2. Molekul mempunyai massa, tetapi
volumenya sangat kecil sehingga dapat
diabaikan
3. Molekul2 berada dalam gerakan acak
konstan pada garis lurus
4. Tabrakan di antara molekul2 dan di antara
molekul dengan dengan dinding pembatas
bersifat elastis yaitu: energi kinetik dapat
ditransfer dari suatu molekul ke molekul
lain, tetapi energi kinetik tersebut tidak
dikonversi ke bentuk energi lain seperti
panas
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Energi Kinetik
Energi Kinetik Translasi (EKT): EKT = ½.m.v2
W = m.g
Dimana: v = kecepatan
w = berat benda
g = gravitasi
Energi Kinetik Rotasi (EKR):
Molekul yang memiliki dua atau lebih atom yang berbeda
(heteronuklir) dapat melakukan gerak berputar
Energi yang timbul akibat rotasi atom dalam molekul
seperti pada peristiwa gelombang mikro spektroskop
disebut: Energi Kinetik Rotasi
Energi Kinetik Vibrasi (EKV):
Energi yang timbul pada pergerakan atom secara vibrasi
dalam molekul disebut: Energi Kinetik Vibrasi.
Dalam peristiwa ini Energi Translasi & Energi Potensial
dipertukarkan secara terus-menerus dan jumlahnya
konstan, asal tidak terjadi pemancaran keluar
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Energi Nuklir
Energi yang dihasilkan dari perubahan status
partikel penyusun inti adalah Energi Nuklir
Energi yang dihasilkan sangat besar reaksi fisi
nuklir atau pembelahan inti (Nuclear Fission)
Contoh:
Uranium (U) yang dibombardir dengan neutron (n)
menghasilkan perubahan jumlah neutron
dalam inti (235 menjadi 236), tetapi jumlah
proton tetap (92)
Bom Hidrogen yang terjadi karena penggabungan
(Fusi) inti Hidrogen
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Energi Elektron
Energi Elektron yaitu: Energi yang timbul karena perubahan
kedudukan orbit elektron
Sesuai dengan model yang diajukan oleh Niels Bohr Atom
mempunyai elektron (e) yang berada pada orbit tertentu dan
bergerak mengitari inti atom
Atom atau molekul mempunyai tingkat energi paling rendah jika
elektron berada pada orbit terendah yang disebut: Orbit Dasar
Gaya yang menahan elektron pada suatu orbit tertentu terdiri atas:
gaya tarik inti ke arah pusat yang dikenal dengan Gaya Coulomb
(gaya centrifugal) sebesar k (Ze2) / r2 dan daya gerak elektron ke
arah luar (gaya centripetal) sebesar mv2/r
Dalam keadaan setimbang, kedua gaya tersebut sama, sehingga
mv2 = k (Ze2) / r2 dimana:
k = konstanta (8,9874 x 109 N.m2/c2)
Z = jumlah proton
e = muatan elektron saat bergerak melintasi garis singgung orbit
Energi elektron semakin kecil (semakin negatif) bila r semakin kecil
sebagai akibat dari energi potensial yang hilang karena elektron
semakin mendekati inti
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Energi Radiasi Matahari
Energi radiasi matahari yang digunakan dalam
proses fotosintesis timbul dari reaksi nuklir yaitu
fusi inti Hidrogen akibat temperatur yang sangat
tinggi
Persamaan Stefan-Boltzmann dapat digunakan
untuk menaksir tingkat radiasi berdasarkan
temperatur: R = εσT4 dimana:
R = pancaran (flux) radiasi (ly/min)
ε = konstanta emisivitas (0 < ε < 1)
= menggambarkan kapasitas suatu
benda mengabsorbsi dan memancarkan
radiasi
σ = konstanta Stefan-Boltzmann
(5,67032 x 10-8 Wm-2K-4)
T = temperatur absolut (273 + 0C)
Radiasi matahari mempunyai suhu pada intinya
sebesar 19.450.000 0C
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Energi Radiasi Matahari
TEORI GELOMBANG:
c=v.λ
Dimana: c = kecepatan cahaya
v = frekuensi
λ = panjang gelombang
Radiasi matahari yang dapat dimanfaatkan oleh tanaman adalah pada
kisaran panjang gelombang 400-700 nm yang disebut: visible
radiation atau Photosynthetically Active Radiation (PAR)
TEORI KUANTA:
Dikembangkan oleh Newton dan Einstein
Merupakan landasan dasar yang digunakan untuk menjelaskan
hubungan energi matahari dengan proses fotosintesis
Partikel energi (Kuanta atau Photon) dari radiasi lebih besar dari
energi yang diperlukan untuk melepaskan elektron dari molekul
elektron akan emisi, dan sebaliknya
Jumlah energi yang dikandung suatu kuanta atau photon
berhubungan dengan panjang gelombang
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Teori Kuanta
Semakin pendek panjang gelombang, maka semakin
tinggi energinya cahaya violet (λ=410 nm)
mempunyai energi lebih besar dari cahaya biru (460
nm), kuning (580 nm), orange (620 nm) dan merah
(660 nm)
E = h.v dimana:
E = energi satu photon cahaya pada
suatu panjang gelombang tertentu
h = konstanta Planck (6,6x10-27erg.det.)
v = frekuensi
Karena: frekuensi = kecepatan cahaya (c) dibagi dengan
panjang gelombang (λ)
maka: E = h.c / λ
Radiasi photon biru dan merah dapat mengeksitasi
elektron molekul fotosintesis (klorofil), tetapi jumlah
maksimum energi yang tersedia biasanya dibatasi
pada panjang gelombang dekat puncak absorpsi
merah (680 nm)
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Teori Kuanta
Energi ekstra dari photon biru tidak dapat digunakan dengan
baik, karena hanya bertahan dalam waktu yang sangat singkat
Energi cahaya merah dengan λ = 680 nm adalah:
E = h.c / λ
E = 6,6x10-27erg.det x 3x1017nm/det x 1/680 nm
= 2,9x10-12erg.det
Catatan:
3x1010cm/det = 3x1017nm/det
1 einstein = 1 mol kuanta
1 photon = 1 kuanta cahaya (radiasi)
1 J.m-2.det-1 = 1 W.m-2
1 cal.cm-2.min-1 = langley = 697,8 W.m-2
10.000 foot-candle = 350 W.m-2
1 foot-candle = 10,76 lux
1 Joule = 107 erg
1 calori = 4,2 Joule
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TUGAS I
TOPIK: REAKSI TERANG FOTOSINTESIS
Oleh: Kelompok 1 Klas L/P/E
Nama Nim.
1.
2. dst
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