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10 BIOLOGY
CHAPTER 2
BIOLOGICAL CLASSIFICATION
2.1 Kingdom Since the dawn of civilisation, there have been many attempts to
Monera
classify living organisms. It was done instinctively not using criteria
2.2 Kingdom Protista that were scientific but borne out of a need to use organisms for our
own use – for food, shelter and clothing. Aristotle was the earliest to
2.3 Kingdom Fungi
attempt a more scientific basis for classification. He used simple
2.4 Kingdom morphological characters to classify plants into trees, shrubs and
Plantae herbs. He also divided animals into two groups, those which had red
2.5 Kingdom blood and those that did not.
Animalia In Linnaeus' time a Two Kingdom system of classification with
Plantae and Animalia kingdoms was developed that included all
2.6 Viruses, Viroids
plants and animals respectively. This system did not distinguish
and Lichens
between the eukaryotes and prokaryotes, unicellular and multicellular
organisms and photosynthetic (green algae) and non-photosynthetic
(fungi) organisms. Classification of organisms into plants and animals
was easily done and was easy to understand, but, a large number of
organisms did not fall into either category. Hence the two kingdom
classification used for a long time was found inadequate. Besides,
gross morphology a need was also felt for including other
characteristics like cell structure, nature of wall, mode of nutrition,
habitat, methods of reproduction, evolutionary relationships, etc.
Classification systems for the living organisms have hence, undergone
several changes over the time. Though plant and animal kingdoms
have been a constant under all different systems, the understanding
of what groups/organisms be included under these kingdoms have
been changing; the number and nature of other kingdoms have also
been understood differently by different scientists over the time.
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BIOLOGICAL CLASSIFICATION 11
Five Kingdoms
Characters
Monera Protista Fungi Plantae Animalia
Cell type Prokaryotic Eukaryotic Eukaryotic Eukaryotic Eukaryotic
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in their walls while the green plants had a cellulosic cell wall. When
such characteristics were considered, the fungi were placed in a
separate kingdom – Kingdom Fungi. All prokaryotic organisms were
grouped together under Kingdom Monera and the unicellular eukaryotic
organisms were placed in Kingdom Protista. Kingdom Protista has
brought together Chlamydomonas, Chlorella (earlier placed in Algae
within Plants and both having cell walls) with Paramoecium and Amoeba
(which were earlier placed in the animal kingdom which lack cell wall). It
has put together organisms which, in earlier classifications, were
placed in different kingdoms. This happened because the criteria for
classification changed. This kind of changes will take place in future
too depending on the improvement in our understanding of
characteristics and evolutionary relationships. Over time, an attempt has
been made to evolve a classification system which reflects not only the
morphological, physiological and reproductive similarities, but is also
phylogenetic, i.e., is based on evolutionary relationships.
In this chapter we will study characteristics of Kingdoms Monera,
Protista and Fungi of the Whittaker system of classification. The
Kingdoms Plantae and Animalia, commonly referred to as plant and
animal kingdoms, respectively, will be dealt separately in chapters 3
and 4.
Spore Flagellum
Cocci Spirilla
Bacilli Vibrio
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BIOLOGICAL CLASSIFICATION 13
2.1.1 Archaebacteria
These bacteria are special since they live in some of the most harsh
habitats such as extreme salty areas (halophiles), hot springs
(thermoacidophiles) and marshy areas (methanogens). Archaebacteria
differ from other bacteria in having a different cell wall structure and this
feature is responsible for their survival in extreme conditions.
Methanogens are present in the gut of several ruminant animals such
as cows and buffaloes and they are responsible for the production of
methane (biogas) from the dung of these animals.
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BIOLOGICAL CLASSIFICATION 15
2.2.1 Chrysophytes
This group includes diatoms and golden algae (desmids). They are
found in fresh water as well as in marine environments. They are
microscopic and float passively in water currents (plankton). Most of
them are photosynthetic. In diatoms the cell walls form two thin
overlapping shells, which fit together as in a soap box. The walls are
embedded with silica and thus the walls are indestructible. Thus,
diatoms have left behind large amount of cell wall deposits in their
habitat; this accumulation over billions of years is referred to as
‘diatomaceous earth’. Being gritty this soil is used in polishing,
filtration of oils and syrups. Diatoms are the chief ‘producers’ in the
oceans.
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2.2.2 Dinoflagellates
These organisms are mostly marine and
photosynthetic. They appear yellow, green, brown, blue
or red depending on the main pigments present in their
cells. The cell wall has stiff cellulose plates on the
outer surface. Most of them have two flagella; one lies
longitudinally and the other transversely in a furrow
between the wall plates. Very often, red
dinoflagellates (Example: Gonyaulax) undergo such
rapid multiplication that they make the sea appear
red (red tides). Toxins released by such large numbers
may even kill other marine animals such as fishes. (a)
2.2.3 Euglenoids
Majority of them are fresh water organisms found in
stagnant water. Instead of a cell wall, they have a
protein rich layer called pellicle which makes their body
flexible. They have two flagella, a short and a long
(b)
one. Though they are photosynthetic in the presence of
sunlight, when deprived of sunlight they behave like
heterotrophs by predating on other smaller organisms.
Interestingly, the pigments of euglenoids are identical
to those present in higher plants. Example: Euglena
(Figure 2.4b).
2.2.5 Protozoans
(d)
All protozoans are heterotrophs and live as predators
or parasites. They are believed to be primitive
Figure 2.4 (a) Dinoflagellates
relatives of animals. There are four major groups of (b) Euglena
protozoans. (c) Slime mould
(d) Paramoecium
Amoeboid protozoans: These organisms live in fresh
water, sea water or moist soil. They move and capture
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BIOLOGICAL CLASSIFICATION 17
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2.3.1 Phycomycetes
Members of phycomycetes are found in aquatic
habitats and on decaying wood in moist and damp
places or as obligate parasites on plants. The (b)
mycelium is aseptate and coenocytic. Asexual
reproduction takes place by zoospores (motile) or by
aplanospores (non-motile). These spores are
endogenously produced in sporangium. A zygospore is
formed by fusion of two gametes. These gametes are
similar in morphology (isogamous) or dissimilar
(anisogamous or oogamous). Some common examples
are Mucor (Figure 2.5a), Rhizopus (the bread mould
mentioned earlier) and Albugo (the parasitic fungi on
mustard).
2.3.2 Ascomycetes
(c)
Commonly known as sac-fungi, the ascomycetes are
mostly multicellular, e.g., Penicillium, or rarely unicellular, Figure 2.5 Fungi: (a) Mucor
e.g., yeast (Saccharomyces). They are saprophytic, (b) Aspergillus (c) Agaricus
decomposers, parasitic or coprophilous (growing on
dung). Mycelium
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BIOLOGICAL CLASSIFICATION 19
2.3.3 Basidiomycetes
Commonly known forms of basidiomycetes are mushrooms, bracket fungi
or puffballs. They grow in soil, on logs and tree stumps and in living
plant bodies as parasites, e.g., rusts and smuts. The mycelium is branched
and septate. The asexual spores are generally not found, but
vegetative reproduction by fragmentation is common. The sex organs
are absent, but plasmogamy is brought about by fusion of two
vegetative or somatic cells of different strains or genotypes. The
resultant structure is dikaryotic which ultimately gives rise to
basidium. Karyogamy and meiosis take place in the basidium
producing four basidiospores. The basidiospores are exogenously
produced on the basidium (pl.: basidia). The basidia are arranged in
fruiting bodies called basidiocarps. Some common members are
Agaricus (mushroom) (Figure 2.5c), Ustilago (smut) and Puccinia (rust
fungus).
2.3.4 Deuteromycetes
Commonly known as imperfect fungi because only the asexual or
vegetative phases of these fungi are known. When the sexual forms of
these fungi were discovered they were moved into classes they rightly
belong to. It is also possible that the asexual and vegetative stage
have been given one name (and placed under deuteromycetes) and the
sexual stage another (and placed under another class). Later when the
linkages were established, the fungi were correctly identified and
moved out of deuteromycetes. Once perfect (sexual) stages of
members of dueteromycetes were discovered they were often moved
to ascomycetes and basidiomycetes. The deuteromycetes reproduce only
by asexual spores known as conidia. The mycelium is septate and
branched. Some members are saprophytes or parasites while a large
number of them are decomposers of litter and help in mineral cycling.
Some examples are Alternaria, Colletotrichum and Trichoderma.
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BIOLOGICAL CLASSIFICATION 21
Head
Collar
Sheath
RNA Capsid
Tail fibres
(a) (b)
Once they infect a cell they take over the machinery of the host cell to
replicate themselves, killing the host. Would you call viruses living or
non-living?
Virus means venom or poisonous fluid. Dmitri Ivanowsky (1892)
recognised certain microbes as causal organism of the mosaic disease
of tobacco (Figure 2.6a). These were found to be smaller than bacteria
because they passed through bacteria-proof filters. M.W. Beijerinek
(1898) demonstrated that the extract of the infected plants of tobacco
could cause infection in healthy plants and named the new pathogen
“virus” and called the fluid as Contagium vivum fluidum (infectious
living fluid). W.M. Stanley (1935) showed that viruses could be
crystallised and crystals consist largely of proteins. They are inert
outside their specific host cell. Viruses are obligate parasites.
In addition to proteins, viruses also contain genetic material, that could
be either RNA or DNA. No virus contains both RNA and DNA. A virus
is a nucleoprotein and the genetic material is infectious. In general,
viruses that infect plants have single stranded RNA and viruses that infect
animals have either single or double stranded RNA or double stranded
DNA. Bacterial viruses or bacteriophages (viruses that infect the
bacteria) are usually double stranded DNA viruses (Figure 2.6b). The
protein coat called capsid made of small subunits called capsomeres,
protects the nucleic acid. These capsomeres are arranged in helical or
polyhedral geometric forms. Viruses cause diseases like mumps,
small pox, herpes and influenza. AIDS in humans is also caused by a
virus. In plants, the symptoms can be mosaic formation, leaf rolling
and curling, yellowing and vein clearing, dwarfing and stunted
growth.
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SUMMARY
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BIOLOGICAL CLASSIFICATION 23
like viruses and viroids as well as the lichens are not included in the five
kingdom system of classification.
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EXERCISES
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