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Porifera phylum
Phylum Porifera
 Porifera means “pore-bearing” and refers to the numerous pores and
channels that permeate a sponge’s body.
 Mostly marine, but include some freshwater inhabitants; usually found
attached to the substratum or objects- occasionally on other animals
such as crabs in shallow or deep water.
 They are sessile; permanently attached to the substrate
 There are approximately 9,000 living species and above 2200 fossil
forms.
 Size – 2 meters to 2 cm
 Saltwater sponges are brightly colored and freshwater sponges are
small and dull green color
 some are round, flat, grow as crusts or vase-like, tube-like shape
 some are radially symmetrical; most are asymmetrical
General Morphology
• The surface of each sponge bears minute pores called ostia (ostium) or
incurrent pores.
•These pores lead into a central hollow cavity, these internal cavity is called
the paragastric cavity or spongocoel
•It opens to outside through a large circular opening, the osculum
• Water is drawn into it through a series of incurrent pores or dermal ostia
present in the body wall into a central cavity and then flows out of the sponge
through a large opening at the top called the osculum.
Body Wall
The thin body wall which encloses the spongocoel, consists of two cellular
layers-
the outer pinacoderm and
inner choanoderm
with a non-cellular mesenchyme in between.
Pinacoderm:
 It is an outer layer of body.
 Thin and flat polygonal cells of pinacoderm is called the pinacocytes
 It protects the internal structure of body
 Each cell presents a central bulging containing a nucleus
 Margins of adjacent cells are closely cemented together
 Pinacocytes are highly contractile, so that the sponge can increase or
decrease slightly in size.

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Obelia life cycle- polyp & medusa structure, difference between polyp & medusa
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Canal system
Canal systemCanal system
Canal system

The canal system in sponges helps with food acquisition, gas exchange, and waste removal. It is composed of incurrent canals that bring water into the body through openings called ostia. The water then flows into radial canals lined with collar cells that have flagella to capture food particles. These radial canals open into a central cavity called the spongocoel. Water then exits through openings called oscula. There are several types of canal systems that vary in complexity, but all involve water entering through ostia, flowing through flagellated chambers that capture food, and exiting through oscula.

 There are special, large and tubular cells, called pore cells and
porocytes
 Each porocyte contains a central canal-like space , these spaces
are called ostia or dermal pores or incurrent pores.
 These permit water to flow from outside into the spongocoel.
 In Syconoid sponge, porocytes connect the incurrent canals
with radial canals
 Through their intracellular channels, the prosopyles
 The openings of incurrent canals into outside are called dermal
ostia and the openings of radial canals into spongocoel are called
apopyles or internal ostia.
Sponge Anatomy
Pechenik, 1996
Pinacocyte
Choanocyte
Amoebocyte
Porocyte
Pinacoderm
Mesenchyme
Choanoderm
Porifera phylum
Choanoderm:
 It is an inner layer of body wall
 A single layer of flagellated collar cells of
choanoderm, are called choanocytes
 A choanocyte is an ovoid cell with its free end
bearing a transparent contractile collar which
surrounds a single long flagellum and a nucleus
at the base or apex of its cell.
 Choanocytes are used in feeding. Collar of
choanocyte traps plankton (food particles) that
are suspended in the water.
 Choanocytes are used in ensuring the flow of
water within the animal’s body by beating their
flagella.
 The water current help in gas exchange,
removal of wastes, and release of the gametes

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Phylum cnidaria
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The phylum Cnidaria includes sea jellies, hydra, coral colonies, sea anemones, sea pens, and sea fans. They are characterized by having radial symmetry, two germ layers, a jelly-like mesoglea layer, an acoelomate body cavity, and specialized stinging cells called cnidocytes. Many species have both a sessile polyp stage and a free-swimming medusa stage. Cnidarians play important ecological roles through food sources, habitat, symbiosis, and wave protection.

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Metagenesis, or alternation of generations, is shown in the life cycle of Obelia sp. The asexual generation is the colonial Obelia polyp, which reproduces asexually through budding. The sexual generation is the solitary medusa, which develops from reproductive polyps in the Obelia colony. The medusa contains gonads and releases gametes that fertilize to form planula larvae. These larvae settle and develop into new Obelia polyp colonies, completing the life cycle through alternating asexual and sexual generations.

Mesenchyme:
 Non-cellular mesenchyme is present between the pinacoderm and the
choanoderm which is secreted by both body layers
 It consists of a gelatinous matrix of protein called mesogloea; contains several
different kinds of freely wandering amoeba-like cells called Amoebocytes
 The minute skeletal elements of mesenchyme is CaCo3 which are called
spicules
 Archaeocytes are amoeboid cells that phagocytize food particles; they can also
undergo differentiation to form other cells, including cells that produce spicules
and gametes
Amoebocytes:
 They may be different types depending upon the shape of their pseudopodia
and function.
 The large-sized primary amoebocytes with blunt pseudopodium and large
nucleus of the cell is called archaeocytes
 These contain much RNA which carry on all the functions essential for the life
of sponge.
 They are self-replicating and also capable of giving rise to all other types of
cells. For this, these cells are called totipotent.
They also give rise to sex cells and play an
important role in regeneration
These cells may be different types-
Scleroblasts: The amoebocytes which manufacture
the skeleton or spicules, are called as scleroblasts.
According to the nature of the product are known as
calcoblasts, silicoblasts or spongioblasts
Collencytes: Most of the amoebocytes have
branching pseudopodia often united into a network
Chromocytes: Pigmented amoebocytes which are
responsible for the color of sponge
Thesocytes: Those amoebocytes which have
rounded pseudopodia and reserve food material.
Myocytes: Those amoebocytes which are present
around osculum.
Gland cells: Those amoebocytes which are attached
to the body surface by long strands and secrete
slime
Sponge Anatomy
Pechenik, 1996
Pinacocyte
Choanocyte
Amoebocyte
Porocyte
Pinacoderm
Mesenchyme
Choanoderm
Porifera phylum

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Skeleton
 The skeleton composed of tiny pointed structures made of
silica or calcium carbonate called spicules which are
embedded in mesenchyme
 Skeleton may also made of network of protein fibers called
Spongin.
 These endoskeleton protects and supports the softer parts of
the body.
 Calcareous spicules are all large spicules.
Types and structure of spicules
Development of spicules
Secreted by specialized
amoebocytes known as
scleroblasts
A monaxon spicule is formed by
two sclerocytes produced by the
division of one scleroblasts
A triaxon spicule is formed by
six sclerocytes produced by the
division of three scleroblasts
Founder cell
Thickener cell
scleroblast
sclerocytes
Water circulation
 The water current used in feeding, gas exchange, removal of wastes, and
release of the gametes
 Because the sponges are sessile
 Water current is caused by constant beating of flagella of choanocytes
 Brings food and oxygen and removes excretory and reproductive elements
Course of water current:
Outside water Ostia Incurrent pore Spongocoel Osculum Outside

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Food and feeding
 Is a filter feeder
 Food consists of planktons (microscopic animals and plants) and organic
particles.
 They get food by circulation of water which enters through the ostia to the
spongocoel.
 Openings are connected by a series of canals, which are lined by
choanocytes (the flagellated collar cells) that maintain the current and filter
out food particles.
 The choanocyte’s collar consists of microvilli joined together by delicate
microfibrils,which filter out tiny food particles.
 The beating of the flagellum draws water through the collar and out the top.
 Particles too big to pass through the collar get trapped in mucus and slide
down the collar to the base where they are phagocytized.
 Some sponges also supplement their filter feeding by hosting symbionts
such as green algae, dinoflagellates or cyanobacteria that provide nutrients
to the sponge.
Canal system
 Inside the body of sponges, the water current flows
through a certain system of spaces collectively forming
the canal system
 Most sponges have one of three types of canal system:
Ascon type
Sycon type
Leucon type
 These systems differ from in each other in the
increasing complexity.
Ascon type
• Simplest type of canal system and found
only in asconoid sponges
• Its have one opening-ostia in its body wall
• The spongocoel is the large and flagellated
cavity
• It is lined by flagellated choanocytes
• Simplest organization,
• Small, branched and tube shaped,
e.g. Leucosolenia, Clathrina canariensis,
Ingressing water
Through
Ostia
spongocoel
Through
Osculum
To outside

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Sycon type
 More complex canal system and found only in
syconoid sponges
 Derived from the asconoid type by horizontal
folding of its wall
 Have two types of canal-incurrent and radial,
paralleling and alternating with each other
 Incurrent canals are non-flagellated and radial
canals are flagellated
 have two openings-dermal ostia and prosopyles
in its body wall
 The spongocoel is the narrow and non-flagellated
cavity
 It is lined by pinacocytes
e.g. Scypha, Grantia
Ingressing water
Dermal
Ostia
Incurrent
canals
Prosopyles
Radial
canals
Apopyles
Spongocoel
Osculum
To outside
Leucon type
• More complex canal system than sycon
type and found only in leuconoid
sponges
• Derived from the sycon type
• Flagellated chambers lined by
choanocytes
• All other spaces are lined by
pinacocytes
• incurrent canals open into Flagellated
cluster chambers through prosopyles
and opens into excurrent canals
through apopyles
• Spongocoel is excessively narrow and
partly disappeared
e.g. Spongilla
Ingressing water
Dermal
Ostia
Incurrent
canals
Prosopyles Flagellated
chambers
Apopyles Excurrent
canals
Osculum
To outside
Class Calcarea (calcareous sponges)
 Small in size and cylindrical and
vase-like in shape
 Only sponges that possess
spicules composed of calcium
carbonate.
 Spicules are straight or have 3-4
rays
 all marine, especially shallow
waters
 Body organization of asconoid,
syconoid and leuconoid type
e.g. Leucosolenia, Scypha
Classification
Class Hexactinellida (Hyalospongiae)
 Moderate sized Glass sponges;
 Body shape cup, urn or vase-like
 characterized by siliceous spicules
 consisting of six rays intersecting at right
angles
 No dermal epithelium or ex-pinacoderm
 Exclusively marine, in deep sea
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1. The document provides an overview of the phylum Cnidaria, which includes jellyfish, corals, and hydroids. 2. Cnidarians are radially or biradially symmetrical and have stinging cells called nematocysts. They exhibit a diploblastic body plan with two tissue layers separated by a gelatinous mesoglea. 3. Many cnidarians alternate between a sessile polyp stage and a motile medusa stage. Reproduction involves external fertilization producing a planula larva.

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The document describes the internal structure and canal system of sponges. It discusses the key components of the canal system including incurrent canals, radial canals, and choanocytes. It describes the four main types of canal systems: Ascon, Sycon, Leucon, and Rhagon. The Ascon type has a simple structure with incurrent pores leading directly to the spongocoel. The Sycon type features incurrent and radial canals. The Leucon type has further folded chambers. The canal systems serve functions like nutrient acquisition, gas exchange, and waste removal for the sponge.

Class Demospongiae (Most sponges)
• Greater than 90 percent of the 5,000
known living sponge species are
demosponges.
• Small to large-sized and vase, cup or
cushion shaped body
• Demosponge skeletons are
composed of spongin fibers or
siliceous spicules or both
• Siliceous spicules with one to four
rays not at right angles,
• All members express the leuconoid
body form
• Mostly found on the continental
shelf in well oxygenated habitats
• All marine except freshwater family
Spongillidae
e.g. Thenea, Spongilla
Yellow sponge growing on a wall
on a Caribbean reef.
Reproduction in Sponges
SEXUAL
• Sponges are monoecious (hermaphrodite) and dioecious
species
• Sperm and ova are derived from archeocytes which
undergo gametogenesis
• Mainly Cross-fertilization occurs because most sponges are
hermaphrodite and produce eggs and sperm at different
times so they do not self fertilize
• Sperms are released to outside through water.
• Choanocytes acting as nurse cells which transport the
sperms to the ova.
• Development takes place within the maternal sponge
• Two types of larvae-
Amphiblastula- characteristics of many calcareous sponges
Parenchymula- characteristics of some calcareous,
Hexactinellida and most Demospongiae sponges
Marine sponges
• Budding
• Fragmentation
• Regeneration
Freshwater
sponges
• Gemmules
• Budding
• Fragmentation
• Regeneration
Porifera phylum
Asexual:
Fragmentation:
When individual pieces
of sponge that are free
floating in the water
column come together
to form a sponge
themselves
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Sponges can grow buds
that become detached
and form sponges
themselves
Gemmules:
Internal buds surrounded by
spongin and spicules that
allow sponges to survive in
harsh conditions; when parent
sponge dies, gemmule
develops into a new sponge
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- Porifera are aquatic animals that live in marine and freshwater environments. They have a simple body plan with pores that allow water flow through a canal system but lack true tissues and organs. - Their cells come in specialized types but can change form, including collar cells that line canals and draw in water and food particles and amoeboid cells that digest and distribute nutrients. - Sponges filter feed but some are predators or parasites. They reproduce asexually through regeneration and buds or sexually with internal fertilization and larval development.

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Sponges are primitive aquatic animals belonging to the phylum Porifera. They come in over 9,000 species and range in size from less than 1 cm to over 1 meter. Sponges have three basic cell types - pinacocytes, mesenchyme cells, and choanocytes. They live in marine environments and have simple structures with no true tissues. Sponges filter feed by drawing water through chambers lined with flagellated choanocytes and trapping particles.

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Invertebrates are animals without backbones. They make up 97% of the animal kingdom and can be found in water and other environments. They exhibit various symmetries and belong to phyla like porifera, cnidaria, platyhelminths, and arthropods. Paramecium are single-celled protozoans that move using cilia, feed by ingesting bacteria and algae, and reproduce through binary fission. Sponges are filter-feeding aquatic animals with pores, skeletal elements, and multiple cell types including choanocytes. Cnidarians like jellyfish have stinging cells and a diploblastic structure, with many exhibiting a complex life cycle involving polyps and medusae

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Porifera phylum

  • 2. Phylum Porifera  Porifera means “pore-bearing” and refers to the numerous pores and channels that permeate a sponge’s body.  Mostly marine, but include some freshwater inhabitants; usually found attached to the substratum or objects- occasionally on other animals such as crabs in shallow or deep water.  They are sessile; permanently attached to the substrate  There are approximately 9,000 living species and above 2200 fossil forms.  Size – 2 meters to 2 cm  Saltwater sponges are brightly colored and freshwater sponges are small and dull green color  some are round, flat, grow as crusts or vase-like, tube-like shape  some are radially symmetrical; most are asymmetrical
  • 3. General Morphology • The surface of each sponge bears minute pores called ostia (ostium) or incurrent pores. •These pores lead into a central hollow cavity, these internal cavity is called the paragastric cavity or spongocoel •It opens to outside through a large circular opening, the osculum • Water is drawn into it through a series of incurrent pores or dermal ostia present in the body wall into a central cavity and then flows out of the sponge through a large opening at the top called the osculum.
  • 4. Body Wall The thin body wall which encloses the spongocoel, consists of two cellular layers- the outer pinacoderm and inner choanoderm with a non-cellular mesenchyme in between. Pinacoderm:  It is an outer layer of body.  Thin and flat polygonal cells of pinacoderm is called the pinacocytes  It protects the internal structure of body  Each cell presents a central bulging containing a nucleus  Margins of adjacent cells are closely cemented together  Pinacocytes are highly contractile, so that the sponge can increase or decrease slightly in size.
  • 5.  There are special, large and tubular cells, called pore cells and porocytes  Each porocyte contains a central canal-like space , these spaces are called ostia or dermal pores or incurrent pores.  These permit water to flow from outside into the spongocoel.  In Syconoid sponge, porocytes connect the incurrent canals with radial canals  Through their intracellular channels, the prosopyles  The openings of incurrent canals into outside are called dermal ostia and the openings of radial canals into spongocoel are called apopyles or internal ostia.
  • 8. Choanoderm:  It is an inner layer of body wall  A single layer of flagellated collar cells of choanoderm, are called choanocytes  A choanocyte is an ovoid cell with its free end bearing a transparent contractile collar which surrounds a single long flagellum and a nucleus at the base or apex of its cell.  Choanocytes are used in feeding. Collar of choanocyte traps plankton (food particles) that are suspended in the water.  Choanocytes are used in ensuring the flow of water within the animal’s body by beating their flagella.  The water current help in gas exchange, removal of wastes, and release of the gametes
  • 9. Mesenchyme:  Non-cellular mesenchyme is present between the pinacoderm and the choanoderm which is secreted by both body layers  It consists of a gelatinous matrix of protein called mesogloea; contains several different kinds of freely wandering amoeba-like cells called Amoebocytes  The minute skeletal elements of mesenchyme is CaCo3 which are called spicules  Archaeocytes are amoeboid cells that phagocytize food particles; they can also undergo differentiation to form other cells, including cells that produce spicules and gametes Amoebocytes:  They may be different types depending upon the shape of their pseudopodia and function.  The large-sized primary amoebocytes with blunt pseudopodium and large nucleus of the cell is called archaeocytes  These contain much RNA which carry on all the functions essential for the life of sponge.  They are self-replicating and also capable of giving rise to all other types of cells. For this, these cells are called totipotent.
  • 10. They also give rise to sex cells and play an important role in regeneration These cells may be different types- Scleroblasts: The amoebocytes which manufacture the skeleton or spicules, are called as scleroblasts. According to the nature of the product are known as calcoblasts, silicoblasts or spongioblasts Collencytes: Most of the amoebocytes have branching pseudopodia often united into a network Chromocytes: Pigmented amoebocytes which are responsible for the color of sponge Thesocytes: Those amoebocytes which have rounded pseudopodia and reserve food material. Myocytes: Those amoebocytes which are present around osculum. Gland cells: Those amoebocytes which are attached to the body surface by long strands and secrete slime
  • 13. Skeleton  The skeleton composed of tiny pointed structures made of silica or calcium carbonate called spicules which are embedded in mesenchyme  Skeleton may also made of network of protein fibers called Spongin.  These endoskeleton protects and supports the softer parts of the body.  Calcareous spicules are all large spicules.
  • 14. Types and structure of spicules
  • 15. Development of spicules Secreted by specialized amoebocytes known as scleroblasts A monaxon spicule is formed by two sclerocytes produced by the division of one scleroblasts A triaxon spicule is formed by six sclerocytes produced by the division of three scleroblasts Founder cell Thickener cell scleroblast sclerocytes
  • 16. Water circulation  The water current used in feeding, gas exchange, removal of wastes, and release of the gametes  Because the sponges are sessile  Water current is caused by constant beating of flagella of choanocytes  Brings food and oxygen and removes excretory and reproductive elements Course of water current: Outside water Ostia Incurrent pore Spongocoel Osculum Outside
  • 18. Food and feeding  Is a filter feeder  Food consists of planktons (microscopic animals and plants) and organic particles.  They get food by circulation of water which enters through the ostia to the spongocoel.  Openings are connected by a series of canals, which are lined by choanocytes (the flagellated collar cells) that maintain the current and filter out food particles.  The choanocyte’s collar consists of microvilli joined together by delicate microfibrils,which filter out tiny food particles.  The beating of the flagellum draws water through the collar and out the top.  Particles too big to pass through the collar get trapped in mucus and slide down the collar to the base where they are phagocytized.  Some sponges also supplement their filter feeding by hosting symbionts such as green algae, dinoflagellates or cyanobacteria that provide nutrients to the sponge.
  • 19. Canal system  Inside the body of sponges, the water current flows through a certain system of spaces collectively forming the canal system  Most sponges have one of three types of canal system: Ascon type Sycon type Leucon type  These systems differ from in each other in the increasing complexity.
  • 20. Ascon type • Simplest type of canal system and found only in asconoid sponges • Its have one opening-ostia in its body wall • The spongocoel is the large and flagellated cavity • It is lined by flagellated choanocytes • Simplest organization, • Small, branched and tube shaped, e.g. Leucosolenia, Clathrina canariensis, Ingressing water Through Ostia spongocoel Through Osculum To outside
  • 21. Sycon type  More complex canal system and found only in syconoid sponges  Derived from the asconoid type by horizontal folding of its wall  Have two types of canal-incurrent and radial, paralleling and alternating with each other  Incurrent canals are non-flagellated and radial canals are flagellated  have two openings-dermal ostia and prosopyles in its body wall  The spongocoel is the narrow and non-flagellated cavity  It is lined by pinacocytes e.g. Scypha, Grantia Ingressing water Dermal Ostia Incurrent canals Prosopyles Radial canals Apopyles Spongocoel Osculum To outside
  • 22. Leucon type • More complex canal system than sycon type and found only in leuconoid sponges • Derived from the sycon type • Flagellated chambers lined by choanocytes • All other spaces are lined by pinacocytes • incurrent canals open into Flagellated cluster chambers through prosopyles and opens into excurrent canals through apopyles • Spongocoel is excessively narrow and partly disappeared e.g. Spongilla Ingressing water Dermal Ostia Incurrent canals Prosopyles Flagellated chambers Apopyles Excurrent canals Osculum To outside
  • 23. Class Calcarea (calcareous sponges)  Small in size and cylindrical and vase-like in shape  Only sponges that possess spicules composed of calcium carbonate.  Spicules are straight or have 3-4 rays  all marine, especially shallow waters  Body organization of asconoid, syconoid and leuconoid type e.g. Leucosolenia, Scypha Classification
  • 24. Class Hexactinellida (Hyalospongiae)  Moderate sized Glass sponges;  Body shape cup, urn or vase-like  characterized by siliceous spicules  consisting of six rays intersecting at right angles  No dermal epithelium or ex-pinacoderm  Exclusively marine, in deep sea E.g. Euplectella (Venus flower)
  • 25. Class Demospongiae (Most sponges) • Greater than 90 percent of the 5,000 known living sponge species are demosponges. • Small to large-sized and vase, cup or cushion shaped body • Demosponge skeletons are composed of spongin fibers or siliceous spicules or both • Siliceous spicules with one to four rays not at right angles, • All members express the leuconoid body form • Mostly found on the continental shelf in well oxygenated habitats • All marine except freshwater family Spongillidae e.g. Thenea, Spongilla Yellow sponge growing on a wall on a Caribbean reef.
  • 26. Reproduction in Sponges SEXUAL • Sponges are monoecious (hermaphrodite) and dioecious species • Sperm and ova are derived from archeocytes which undergo gametogenesis • Mainly Cross-fertilization occurs because most sponges are hermaphrodite and produce eggs and sperm at different times so they do not self fertilize • Sperms are released to outside through water. • Choanocytes acting as nurse cells which transport the sperms to the ova. • Development takes place within the maternal sponge • Two types of larvae- Amphiblastula- characteristics of many calcareous sponges Parenchymula- characteristics of some calcareous, Hexactinellida and most Demospongiae sponges Marine sponges • Budding • Fragmentation • Regeneration Freshwater sponges • Gemmules • Budding • Fragmentation • Regeneration
  • 28. Asexual: Fragmentation: When individual pieces of sponge that are free floating in the water column come together to form a sponge themselves Budding: Sponges can grow buds that become detached and form sponges themselves Gemmules: Internal buds surrounded by spongin and spicules that allow sponges to survive in harsh conditions; when parent sponge dies, gemmule develops into a new sponge when conditions are favorable